• A History of the Ferret

    Item Type Journal Article
    Author ALEXANDER P. D. THOMSON
    Date 1951-10-01
    Library Catalog Silverchair
    URL https://doi.org/10.1093/jhmas/VI.Autumn.471
    Accessed 2/20/2025, 6:00:45 PM
    Volume VI
    Pages 471-480
    Publication Journal of the History of Medicine and Allied Sciences
    DOI 10.1093/jhmas/VI.Autumn.471
    Issue Autumn
    Journal Abbr Journal of the History of Medicine and Allied Sciences
    ISSN 0022-5045
    Date Added 2/20/2025, 6:00:45 PM
    Modified 10/23/2025, 10:25:39 AM

    Attachments

    • PDF
    • Snapshot
  • A new invasive species in the Canary Islands: a naturalized population of ferrets Mustela furo in La Palma Biosphere Reserve

    Item Type Journal Article
    Author Félix M. Medina
    Author Aurelio Martín
    Abstract Abstract We present data to confirm that the island of La Palma harbours the first established feral population of ferrets Mustela furo on the Canary Islands in the Atlantic Ocean. It is not known when the species was introduced but individuals occasionally lost during hunting appear to have become established sometime in the previous 2 decades. Sightings of ferrets are mainly in the north of the island but they are likely to expand their range southwards, and a few have already been detected in the centre of the island. We report a total of 45 cases of wild ferrets in 28 different localities during 1998-2007. To minimize effects on native species, control measures or eradication are required. Public awareness of the problem and education campaigns, especially among hunters, are needed to reduce the threat of this alien species to the biota of the Canary Islands. [PUBLICATION ABSTRACT]
    Date Jan 2010
    Language English
    Short Title A new invasive species in the Canary Islands
    Library Catalog ProQuest
    URL https://search.proquest.com/docview/222308584/abstract/384EDBD14684498BPQ/1
    Accessed 4/29/2019, 10:57:11 AM
    Rights Copyright © Fauna & Flora International 2009
    Volume 44
    Pages 41-44
    Publication Oryx; Cambridge
    DOI http://dx.doi.org/10.1017/S0030605309990743
    Issue 1
    ISSN 00306053
    Date Added 4/29/2019, 10:57:11 AM
    Modified 5/21/2025, 4:36:49 PM

    Tags:

    • Conservation
    • Biology--Zoology

    Attachments

    • Medina and Martín - 2010 - A new invasive species in the Canary Islands a na.pdf
  • A novel A (H7N2) influenza virus isolated from a veterinarian caring for cats in a New York City animal shelter causes mild disease and transmits poorly in the ferret model

    Item Type Journal Article
    Author Jessica A. Belser
    Author Joanna A. Pulit-Penaloza
    Author Xiangjie Sun
    Author Nicole Brock
    Author Claudia Pappas
    Author Hannah M. Creager
    Author Hui Zeng
    Author Terrence M. Tumpey
    Author Taronna R. Maines
    Abstract In December 2016, a low-pathogenic avian influenza (LPAI) A(H7N2) virus was identified to be the causative source of an outbreak in a cat shelter in New York City, which subsequently spread to multiple shelters in the states of New York and Pennsylvania. One person with occupational exposure to infected cats became infected with the virus, representing the first LPAI H7N2 virus infection in a human in North America since 2003. Considering the close contact that frequently occurs between companion animals and humans, it was critical to assess the relative risk of this novel virus to public health. The virus isolated from the human case, A/New York/108/2016 (NY/108), caused mild and transient illness in ferrets and mice but did not transmit to naive cohoused ferrets following traditional or aerosol-based inoculation methods. The environmental persistence of NY/108 virus was generally comparable to that of other LPAI H7N2 viruses. However, NY/108 virus replicated in human bronchial epithelial cells with an increased efficiency compared with that of previously isolated H7N2 viruses. Furthermore, the novel H7N2 virus was found to utilize a relatively lower pH for hemagglutinin activation, similar to human influenza viruses. Our data suggest that the LPAI H7N2 virus requires further adaptation before representing a substantial threat to public health. However, the reemergence of an LPAI H7N2 virus in the northeastern United States underscores the need for continuous surveillance of emerging zoonotic influenza viruses inclusive of mammalian species, such as domestic felines, that are not commonly considered intermediate hosts for avian influenza viruses.IMPORTANCE Avian influenza viruses are capable of crossing the species barrier to infect mammals, an event of public health concern due to the potential acquisition of a pandemic phenotype. In December 2016, an H7N2 virus caused an outbreak in cats in multiple animal shelters in New York State. This was the first detection of this virus in the northeastern United States in over a decade and the first documented infection of a felid with an H7N2 virus. A veterinarian became infected following occupational exposure to H7N2 virus-infected cats, necessitating the evaluation of this virus for its capacity to cause disease in mammals. While the H7N2 virus was associated with mild illness in mice and ferrets and did not spread well between ferrets, it nonetheless possessed several markers of virulence for mammals. These data highlight the promiscuity of influenza viruses and the need for diligent surveillance across multiple species to quickly identify an emerging strain with pandemic potential.
    Date 2017-07-12
    Language en
    Library Catalog journals.asm.org (Atypon)
    URL https://journals.asm.org/doi/full/10.1128/jvi.00672-17
    Accessed 10/22/2024, 4:49:07 PM
    Extra Publisher: American Society for Microbiology
    Volume 91
    Pages 10.1128/jvi.00672-17
    Publication Journal of Virology
    DOI 10.1128/jvi.00672-17
    Issue 15
    Journal Abbr J Virol
    ISSN 0022-538X, 1098-5514
    Date Added 7/14/2023, 12:18:35 PM
    Modified 5/21/2025, 4:36:30 PM

    Attachments

    • Full Text PDF
  • A panel of microsatellite markers for genetic studies of European polecats (Mustela putorius) and ferrets (Mustela furo)

    Item Type Journal Article
    Author Mafalda Costa
    Author Carlos Fernandes
    Author Mónica Rodrigues
    Author Margarida Santos-Reis
    Author Michael W. Bruford
    Abstract We report a panel of 12 microsatellite markers, including nine novel polymorphic loci isolated in the European polecat and three loci previously developed in closely related species, for genetic studies of polecats and ferrets. We tested the panel at fine geographic scales in polecat and domestic ferret populations of Britain and Portugal and at a broad geographic scale, using 50 polecat samples from across Western Europe. At the fine geographic scales, we recorded one to five alleles per locus and observed and expected heterozygosities ranging from 0 to 0.80 and from 0 to 0.74, respectively. These values increased markedly in the European sample of polecats. No linkage disequilibrium was detected between any pair of loci and all genotypic frequencies complied with Hardy–Weinberg equilibrium expectations. The microsatellite panel allowed the molecular discrimination of polecats and domestic ferrets in Britain, where hybridisation between the two species has been documented. Thus, these loci may be useful not only in genetic studies of polecats and ferrets but also in studies to assess hybridisation between these two mustelid species.
    Date 2012-06-01
    Language en
    Library Catalog Springer Link
    URL https://doi.org/10.1007/s10344-012-0627-1
    Accessed 5/21/2025, 4:02:33 PM
    Volume 58
    Pages 629-633
    Publication European Journal of Wildlife Research
    DOI 10.1007/s10344-012-0627-1
    Issue 3
    Journal Abbr Eur J Wildl Res
    ISSN 1439-0574
    Date Added 5/21/2025, 4:02:34 PM
    Modified 5/22/2025, 8:07:08 PM

    Tags:

    • Microsatellites
    • Mustela putorius
    • European polecat
    • Ferret
    • Genetic mapping
    • Genetic Markers
    • Genotyping and haplotyping
    • Hybridisation
    • Microsatellite instability
    • Mustela furo
    • Population Genetics
    • Quantitative trait loci

    Attachments

    • Costa et al. - 2012 - A panel of microsatellite markers for genetic studies of European polecats (Mustela putorius) and fe.pdf
  • A rating system for potential exotic bird and mammal pests

    Item Type Journal Article
    Author K. Shawn Smallwood
    Author Terrell P. Salmon
    Date 1992
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://linkinghub.elsevier.com/retrieve/pii/000632079291043R
    Accessed 7/14/2023, 1:09:04 PM
    Rights https://www.elsevier.com/tdm/userlicense/1.0/
    Volume 62
    Pages 149-159
    Publication Biological Conservation
    DOI 10.1016/0006-3207(92)91043-R
    Issue 3
    Journal Abbr Biological Conservation
    ISSN 00063207
    Date Added 7/14/2023, 1:09:04 PM
    Modified 5/21/2025, 4:37:20 PM

    Attachments

    • Shawn Smallwood and Salmon - 1992 - A rating system for potential exotic bird and mammal pests.pdf
  • A review of national and California population estimates of pet ferrets

    Item Type Report
    Author Ronald M Jurek
    Abstract The number of ferrets illegally owned in California has never been properly assessed, yet some pet ferret groups have routinely cited estimates of this animal’s abundance in discussions about matters of legislation, wildlife conservation, and public health concerns. Since the mid 1980s, organizations promoting the legalization of ferrets for pet keeping have claimed that hundreds of thousands of illegally owned ferrets are already in California. Some groups have portrayed domestic ferrets as the “Pet of the 90s,” and with numbers approaching 10 million in the United States, they claim, ferrets have now become the third most numerous or popular companion animal (or “interactive” pet) in the nation. Some ferret groups have claimed that more than a half million, and up to one million, ferrets are illegally owned in California. However, nationwide pet ownership surveys do not support these claims, indicating that approximately 800,000 pet ferrets existed in the entire nation in 1996. This implies, therefore, that in recent years, there have been fewer than--possibly far fewer than--l00,000 illegally owned ferrets in California.
    Date 1998
    Language en
    Library Catalog Zotero
    Place Sacramento, CA
    Report Number 98-09
    Report Type Bird and Mammal Conservation Program Report
    Institution California Department Fish Wildlife
    Date Added 3/25/2025, 11:16:52 AM
    Modified 10/21/2025, 3:51:33 PM

    Attachments

    • Jurek - A REVIEW OF NATIONAL AND CALIFORNIA POPULATION ESTIMATES OF PET FERRETS.pdf
  • A simple model for ferret population dynamics and control in semi-arid New Zealand habitats

    Item Type Journal Article
    Author N. D. Barlow
    Author G. L. Norbury
    Abstract Introduced ferrets (Mustela furo) in New Zealand are subject to population control to reduce their threat to native fauna and the incidence of bovine tuberculosis (Tb) in livestock. To help in evaluating control options and to contribute to a multi-species model for Tb dynamics, a simple Ricker model was developed for ferret population dynamics in a semi-arid environment. The model was based on two data sets and suggested an intrinsic rate of increase for ferrets of 1.0–1.3 year–1 and a carrying capacity of 0.5–2.9 km–2. There was evidence for direct density-dependence in both data sets and the effect appeared to act mainly on recruitment. Dependence of the rate of increase of predators on the density of wild rabbits (Oryctolagus cuniculus) was exhibited in one of the two data sets, together with a numerical response relating current density of predators asymptotically to current density of rabbits, their primary prey. Predators in this data set included both cats and ferrets, estimated from spotlight counts, but the other data set demonstrated a direct proportionality between predator (cat and ferret) spotlight counts and minimum ferrets known to be alive by trapping. The model suggested, firstly, that populations are hard to suppress by continuous culling, with at least a 50% removal per year necessary to effect a suppression of 50% in long-term average density. Secondly, if control is episodic rather than continuous, culling in autumn gives a greater degree of suppression over time (280%, accumulated over time) than culling in spring (180%). A differential equation version of the model provides a component for a general Anderson/May bovine Tb/wildlife (possum/deer/ferret) model.
    Date 2001
    Language en
    Library Catalog www.publish.csiro.au
    URL http://www.publish.csiro.au/wr/WR99090
    Accessed 4/29/2019, 10:47:14 AM
    Volume 28
    Pages 87-94
    Publication Wildlife Research
    DOI 10.1071/wr99090
    Issue 1
    Journal Abbr Wildl. Res.
    ISSN 1448-5494
    Date Added 4/29/2019, 10:47:14 AM
    Modified 5/21/2025, 4:37:51 PM

    Attachments

    • Barlow and Norbury - 2001 - A simple model for ferret population dynamics and control in semi-arid New Zealand habitats.pdf
    • Snapshot
  • Advances in New Zealand mammalogy 1990–2000: Feral ferret

    Item Type Journal Article
    Author B. K. Clapperton
    Date 03/2001
    Language en
    Short Title Advances in New Zealand mammalogy 1990–2000
    Library Catalog DOI.org (Crossref)
    URL http://www.tandfonline.com/doi/abs/10.1080/03014223.2001.9517647
    Accessed 4/29/2019, 10:50:11 AM
    Volume 31
    Pages 185-203
    Publication Journal of the Royal Society of New Zealand
    DOI 10.1080/03014223.2001.9517647
    Issue 1
    Journal Abbr Journal of the Royal Society of New Zealand
    ISSN 0303-6758, 1175-8899
    Date Added 4/29/2019, 10:50:11 AM
    Modified 5/21/2025, 4:38:15 PM

    Attachments

    • Clapperton - 2001 - Advances in New Zealand mammalogy 1990–2000 Feral ferret.pdf
  • Analysis of the potential impacts of domesticated ferrets upon wildlife, agriculture, and human health in North America, with a focus upon California, based upon literature review and survey of North American governmental agencies

    Item Type Report
    Author Gary Orval Graening
    Date 9/13/2010
    Place Sacramento, CA
    Institution California State University
    Date Added 7/14/2023, 12:14:28 PM
    Modified 10/22/2024, 4:53:00 PM

    Tags:

    • KEY PAPER

    Attachments

    • Final_Report_unlocked.pdf
  • Assessing the host disease status of wildlife and the implications for disease control: Mycobacterium bovis infection in feral ferrets

    Item Type Journal Article
    Author Peter Caley
    Author Jim Hone
    Date 2005-06-01
    Language en
    Short Title Assessing the host disease status of wildlife and the implications for disease control
    Library Catalog DOI.org (Crossref)
    URL http://doi.wiley.com/10.1111/j.1365-2664.2005.01053.x
    Accessed 4/29/2019, 10:48:47 AM
    Rights http://onlinelibrary.wiley.com/termsAndConditions#vor
    Volume 42
    Pages 708-719
    Publication Journal of Applied Ecology
    DOI 10.1111/j.1365-2664.2005.01053.x
    Issue 4
    ISSN 00218901, 13652664
    Date Added 4/29/2019, 10:48:47 AM
    Modified 5/21/2025, 4:38:40 PM

    Attachments

    • Caley and Hone - 2005 - Assessing the host disease status of wildlife and .pdf
  • Bayesian analysis of hybridization and introgression between the endangered european mink (Mustela lutreola) and the polecat (Mustela putorius)

    Item Type Journal Article
    Author M. T. Cabria
    Author J. R. Michaux
    Author B. J. Gómez-Moliner
    Author D. Skumatov
    Author T. Maran
    Author P. Fournier
    Author J. López de Luzuriaga
    Author R. Zardoya
    Abstract Human-mediated global change will probably increase the rates of natural hybridization and genetic introgression between closely related species, and this will have major implications for conservation of the taxa involved. In this study, we analyse both mitochondrial and nuclear data to characterize ongoing hybridization and genetic introgression between two sympatric sister species of mustelids, the endangered European mink (Mustela lutreola) and the more abundant polecat (M. putorius). A total of 317 European mink, 114 polecats and 15 putative hybrid individuals were collected from different localities in Europe and genotyped with 13 microsatellite nuclear markers. Recently developed Bayesian methods for assigning individuals to populations and identifying admixture proportions were applied to the genetic data. To identify the direction of hybridization, we additionally sequenced mtDNA and Y chromosomes from 78 individuals and 29 males respectively. We found that both hybridization and genetic introgression occurred at low levels (3% and 0.9% respectively) and indicated that hybridization is asymmetric, as only pure polecat males mate with pure European mink females. Furthermore, backcrossing and genetic introgression was detected only from female first-generation (F1) hybrids of European mink to polecats. This latter result implies that Haldane's rule may apply. Our results suggest that hybridization and genetic introgression between the two species should be considered a rather uncommon event. However, the current low densities of European mink might be changing this trend.
    Date Mar 2011
    Language eng
    Library Catalog PubMed
    URL https://onlinelibrary.wiley.com/doi/10.1111/j.1365-294X.2010.04988.x
    Rights http://onlinelibrary.wiley.com/termsAndConditions#vor
    Extra PMID: 21244536
    Volume 20
    Pages 1176-1190
    Publication Molecular Ecology
    DOI 10.1111/j.1365-294X.2010.04988.x
    Issue 6
    Journal Abbr Mol. Ecol.
    ISSN 1365-294X
    Date Added 4/29/2019, 10:58:27 AM
    Modified 5/21/2025, 4:47:46 PM

    Tags:

    • Animals
    • Male
    • Bayes Theorem
    • DNA, Mitochondrial
    • Mink
    • Ferrets
    • Female
    • Microsatellite Repeats
    • Hybridization, Genetic
    • Molecular Sequence Data
    • Y Chromosome

    Attachments

    • Cabria et al. - 2011 - Bayesian analysis of hybridization and introgressi.pdf
    • PubMed entry
  • Black-footed ferret (Mustela nigripes) population dynamics: use of simulation modeling to investigate the effects of stochasticity, habitat geometry and dispersal

    Item Type Thesis
    Author Robert W. Van Kirk
    Date 1990
    Language en
    Short Title Black-footed ferret(Mustela nigripes) population dynamics
    Library Catalog Google Scholar
    URL https://humboldt-dspace.calstate.edu/handle/2148/878
    Accessed 2/10/2014, 6:59:46 AM
    Place Arcata, CA
    # of Pages 97
    Type Master's Thesis
    University Humboldt State University
    Date Added 2/10/2014, 6:59:46 AM
    Modified 5/21/2025, 4:48:45 PM

    Attachments

    • Van Kirk - 1990 - Black-footed ferret (Mustela nigripes) population .pdf
  • California law and ferrets: are they truly wild weasels?

    Item Type Journal Article
    Author David L Herman
    Date 2000
    Language en
    Library Catalog Zotero
    Volume 23
    Pages 37-53
    Publication Environs
    Issue 2
    Date Added 2/13/2023, 7:58:59 AM
    Modified 5/21/2025, 4:49:14 PM

    Attachments

    • Herman - 2000 - California law and ferrets are they truly wild we.pdf
  • Canine distemper in black-footed ferrets (Mustela nigripes) from Wyoming

    Item Type Journal Article
    Author E. S. Williams
    Author E. T. Thome
    Author M. J. G. Appel
    Author D. W. Belitsky
    Abstract In September and October 1985, six black-footed ferrets (Mustela ntgripes) were captured from the only known population, located near Meeteetse, Wyoming for captive propagation. Two days following capture an adult male showed signs of canine distemper and an adult female displayed similar signs 7 days postcapture; these infections were undoubtedly acquired prior to capture. Subsequently the four remaining captive black-footed ferrets also developed canine distemper and all eventually died. Clinical signs included severe pruritus, hyperkeratosis and progressive loss of body condition. A few animals had intermittent diarrhea and respiratory disease. Intranuclear and intracytoplasmic inclusion bodies were numerous in epithelial tissues and two black-footed ferrets had a mild to moderate meningoencephalitis. Canine distemper virus was isolated from four animals and paramyxovirus nucleocapsids were observed by electron microscopy of feces from all affected black-footed ferrets. Antibodies to canine distemper virus were not detected in sera of sick black-footed ferrets. Antibodies to canine distemper virus were found in sera of badgers (Taxidea taxus) and coyotes (Canis latrans) collected in the Meeteetse area in 1986. Most free-ranging black-footed ferrets in the colony apparently died of canine distemper during the summer and fall of 1985. An attempt was made to capture all surviving animals in the affected area in order to abort the epizootic and provide black-footed ferrets for captive propagation.
    Date 07/1988
    Language en
    Library Catalog Crossref
    URL http://www.jwildlifedis.org/doi/10.7589/0090-3558-24.3.385
    Accessed 8/7/2018, 3:23:36 PM
    Volume 24
    Pages 385-398
    Publication Journal of Wildlife Diseases
    DOI 10.7589/0090-3558-24.3.385
    Issue 3
    ISSN 0090-3558
    Date Added 8/7/2018, 3:23:36 PM
    Modified 5/21/2025, 4:49:35 PM

    Attachments

    • Williams et al. - 1988 - Canine distemper in black-footed ferrets (Mustela .pdf
  • Comparison of black-footed ferret (Mustela nigripes) and domestic ferret (M. putorius furo) courtship activity

    Item Type Journal Article
    Author Brian J. Miller
    Author Stanley H. Anderson
    Abstract Because of the scarcity of the endangered black-footed ferret (Mustela nigripes) and the amount of knowledge necessary for their conservation, surrogate research can play an important role in recovery. In this paper, we investigate surrogate behavioral research potential by comparing courtship behavior of the black-footed ferret to the congeneric domestic ferret (M. putorius furo). Ten female domestic ferrets were bred to five male domestic ferrets and eight female black-footed ferrets were bred to five black-footed ferret males. Courtship activities were defined, analyzed, and quantitatively compared between both groups. Lag sequential analysis of was used to prepare the behavioral matrices, and matrix cells were compared between groups with an equality of proportions test. Courtship patterns did not differ significantly between the two closely related species, and the domestic ferret would probably be an adequate surrogate for reproductive behavior research on the black-footed ferret.
    Date 1990
    Language en
    Library Catalog Wiley Online Library
    URL https://onlinelibrary.wiley.com/doi/abs/10.1002/zoo.1430090303
    Accessed 2/20/2025, 6:01:27 PM
    Rights Copyright © 1990 Wiley-Liss, Inc., A Wiley Company
    Extra _eprint: https://onlinelibrary.wiley.com/doi/pdf/10.1002/zoo.1430090303
    Volume 9
    Pages 201-210
    Publication Zoo Biology
    DOI 10.1002/zoo.1430090303
    Issue 3
    ISSN 1098-2361
    Date Added 2/20/2025, 6:01:27 PM
    Modified 10/23/2025, 10:27:34 AM

    Tags:

    • ferret behavior
    • pre-couplatory behavior
    • reproductive behavior

    Attachments

    • PDF
    • Snapshot
  • Daily activity of stoats (Mustela erminea), feral ferrets (Mustela furo) and feral house cats (Felis catus) in coastal grassland, Otago Peninsula, New Zealand.

    Item Type Journal Article
    Author N Alterio
    Author H Moller
    Date 1997
    Language en
    Library Catalog Zotero
    Volume 21
    Pages 89-95
    Publication New Zealand Journal of Ecology
    Issue 1
    Date Added 4/29/2019, 10:54:38 AM
    Modified 5/21/2025, 4:50:14 PM

    Attachments

    • Alterio and Moller - 1997 - Daily activity of stoats (Mustela erminea), feral .pdf
  • Dispersal and survival of juvenile feral ferrets Mustela furo in New Zealand

    Item Type Journal Article
    Author Andrea E. Byrom
    Date 02/2002
    Language en
    Library Catalog DOI.org (Crossref)
    URL http://doi.wiley.com/10.1046/j.1365-2664.2002.00689.x
    Accessed 4/29/2019, 10:53:28 AM
    Rights http://onlinelibrary.wiley.com/termsAndConditions#vor
    Volume 39
    Pages 67-78
    Publication Journal of Applied Ecology
    DOI 10.1046/j.1365-2664.2002.00689.x
    Issue 1
    Journal Abbr J Appl Ecology
    ISSN 0021-8901, 1365-2664
    Date Added 4/29/2019, 10:53:28 AM
    Modified 5/21/2025, 4:50:29 PM

    Attachments

    • Byrom - 2002 - Dispersal and survival of juvenile feral ferrets M.pdf
  • Domestication effect of reduced brain size is reverted when mink become feral

    Item Type Journal Article
    Author Ann-Kathrin Pohle
    Author Andrzej Zalewski
    Author Marion Muturi
    Author Christian Dullin
    Author Lucie Farková
    Author Lara Keicher
    Author Dina K. N. Dechmann
    Abstract A typical consequence of breeding animal species for domestication is a reduction in relative brain size. When domesticated animals escape from captivity and establish feral populations, the larger brain of the wild phenotype is usually not regained. In the American mink (Neovison vison), we found an exception to this rule. We confirmed the previously described reduction in relative braincase size and volume compared to their wild North American ancestors in mink bred for their fur in Poland, in a dataset of 292 skulls. We then also found a significant regrowth of these measures in well-established feral populations in Poland. Closely related, small mustelids are known for seasonal reversible changes in skull and brain size. It seems that these small mustelids are able to regain the brain size, which is adaptive for living in the wild, and flexibly respond to selection accordingly.
    Date 2023-07-05
    Library Catalog royalsocietypublishing.org (Atypon)
    URL https://royalsocietypublishing.org/doi/10.1098/rsos.230463
    Accessed 7/14/2025, 4:00:23 PM
    Extra Publisher: Royal Society
    Volume 10
    Pages 230463
    Publication Royal Society Open Science
    DOI 10.1098/rsos.230463
    Issue 7
    Date Added 7/14/2025, 4:00:23 PM
    Modified 7/14/2025, 4:00:23 PM

    Tags:

    • brain size
    • domestication
    • feralization
    • American mink

    Attachments

    • Full Text PDF
  • Ecology and conservation of the polecat Mustela putorius (Linnaeus, 1758) in Portugal: a review

    Item Type Journal Article
    Author Mafalda Costa
    Author Carlos Fernandes
    Author Margarida Santos-Reis
    Date 2014
    Short Title Ecology and conservation of the polecat Mustela putorius (Linnaeus, 1758) in Portugal
    Library Catalog Google Scholar
    URL https://www.aranzadi.eus/fileadmin/docs/Munibe/2014079087.pdf
    Accessed 10/22/2025, 7:52:48 AM
    Volume 3
    Pages 79–87
    Publication Munibe Monographs Nature Series
    Date Added 10/22/2025, 7:52:48 AM
    Modified 10/22/2025, 7:52:48 AM

    Attachments

    • Available Version (via Google Scholar)
  • Effect of captivity and management on behaviour of the domestic ferret (Mustela putorius furo)

    Item Type Journal Article
    Author Sarah Talbot
    Author Rafael Freire
    Author Skye Wassens
    Date 02/2014
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://linkinghub.elsevier.com/retrieve/pii/S016815911300289X
    Accessed 5/21/2025, 4:10:25 PM
    Volume 151
    Pages 94-101
    Publication Applied Animal Behaviour Science
    DOI 10.1016/j.applanim.2013.11.017
    Journal Abbr Applied Animal Behaviour Science
    ISSN 01681591
    Date Added 5/21/2025, 4:10:25 PM
    Modified 5/21/2025, 4:10:25 PM

    Attachments

    • Talbot et al. - 2014 - Effect of captivity and management on behaviour of the domestic ferret (Mustela putorius furo).pdf
  • Environmental impact and relative invasiveness of free-roaming domestic carnivores—a North American survey of governmental agencies

    Item Type Journal Article
    Author Ana Lepe
    Author Valerie Kaplan
    Author Alirio Arreaza
    Author Robert Szpanderfer
    Author David Bristol
    Author M. Scott Sinclair
    Abstract Simple Summary This paper reports on a survey that explores the impact of three non-native domestic carnivores—dogs, cats, and ferrets—on the native wildlife of the United States (US) and Canada. Government agencies were asked to document the number and frequency of sightings, and the degree of concern resulting from free-roaming animals on urban, suburban, rural, recreational areas, and wildlands in their jurisdictions. Results confirm the existence of free-roaming cats and dogs throughout North America, as well as their profound impact on native wildlife, with cats being the major offenders. Except for an occasional stray, free-roaming ferrets were “never” or “rarely seen”; no agency reported that ferrets caused environmental harm. This is the first study to compare the relative impact of free-roaming dogs, cats, and ferrets. It shows differences in how these three animals react to novel environments. For the US and Canada, free roaming cats and dogs meet the definition of an “invasive” species, whereas ferrets do not. However, the way we as a society view these animals, our attitudes and perceptions, may influence how governmental agencies manage and control them. Abstract A survey of the United States and Canadian governmental agencies investigated the environmental impact and relative invasiveness of free-roaming domestic non-native carnivores—dogs, cats, and ferrets. Agencies represented wildlife, fish, game, natural or environmental resources, parks and recreation, veterinary and human health, animal control, and agriculture. Respondents were asked to document the number and frequency of sightings of unconfined animals, evidence for environmental harm, and the resulting “degree of concern” in their respective jurisdictions. Results confirmed the existence of feral (breeding) cats and dogs, documenting high levels of concern regarding the impact of these animals on both continental and surrounding insular habitats. Except for occasional strays, no free-roaming or feral ferrets were reported; nor were there reports of ferrets impacting native wildlife, including ground-nesting birds, or sensitive species. This is the first study to report the relative impact of free-roaming domestic carnivores. Dogs and cats meet the current definition of “invasive” species, whereas ferrets do not. Differences in how each species impacts the North American environment highlights the complex interaction between non-native species and their environment. Public attitudes and perceptions regarding these species may be a factor in their control and agency management priorities.
    Date 2017-10-14
    Language en
    Library Catalog PubMed Central
    URL https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5664037/
    Accessed 7/14/2023, 7:10:32 PM
    Rights https://creativecommons.org/licenses/by/4.0/
    Extra PMID: 29036923 PMCID: PMC5664037
    Volume 7
    Pages 78
    Publication Animals : an Open Access Journal from MDPI
    DOI 10.3390/ani7100078
    Issue 10
    Journal Abbr Animals (Basel)
    ISSN 2076-2615
    Date Added 7/14/2023, 7:10:32 PM
    Modified 8/20/2025, 11:10:06 AM

    Tags:

    • invasive
    • North America
    • non-native species
    • government
    • dog
    • survey
    • environmental impact
    • ferret
    • cat
    • domestic carnivore

    Attachments

    • Lepe et al. - 2017 - Environmental impact and relative invasiveness of .pdf
    • PubMed Central Link
  • Estimating the force of infection; Mycobacterium bovis infection in feral ferrets Mustela furo in New Zealand

    Item Type Journal Article
    Author Peter Caley
    Author Jim Hone
    Date 01/2002
    Language en
    Library Catalog DOI.org (Crossref)
    URL http://doi.wiley.com/10.1046/j.0021-8790.2001.00573.x
    Accessed 4/29/2019, 10:49:20 AM
    Rights http://onlinelibrary.wiley.com/termsAndConditions#vor
    Volume 71
    Pages 44-54
    Publication Journal of Animal Ecology
    DOI 10.1046/j.0021-8790.2001.00573.x
    Issue 1
    Journal Abbr J Anim Ecology
    ISSN 0021-8790, 1365-2656
    Date Added 4/29/2019, 10:49:20 AM
    Modified 5/21/2025, 4:51:34 PM

    Attachments

    • Caley and Hone - 2002 - Estimating the force of infection; Mycobacterium b.pdf
  • Extensive genome introgression between domestic ferret and European polecat during population recovery in Great Britain

    Item Type Journal Article
    Author Graham J Etherington
    Author Adam Ciezarek
    Author Rebecca Shaw
    Author Johan Michaux
    Author Elizabeth Croose
    Author Wilfried Haerty
    Author Federica Di Palma
    Editor Liliana Cortés-Ortiz
    Abstract The European polecat (Mustela putorius) is a mammalian predator which occurs across much of Europe east to the Ural Mountains. In Great Britain, following years of persecution the range of the European polecat contracted and by the early 1900s was restricted to unmanaged forests of central Wales. The European polecat has recently undergone a population increase due to legal protection and its range now overlaps that of feral domestic ferrets (Mustela putorius furo). During this range expansion, European polecats hybridized with feral domestic ferrets producing viable offspring. Here, we carry out population-level whole-genome sequencing on 8 domestic ferrets, 19 British European polecats, and 15 European polecats from the European mainland. We used a range of population genomics methods to examine the data, including phylogenetics, phylogenetic graphs, model-based clustering, phylogenetic invariants, ABBA-BABA tests, topology weighting, and Fst. We found high degrees of genome introgression in British polecats outside their previous stronghold, even in those individuals phenotyped as “pure” polecats. These polecats ranged from presumed F1 hybrids (gamma = 0.53) to individuals that were much less introgressed (gamma = 0.2). We quantify this introgression and find introgressed genes containing Fst outliers associated with cognitive function and sight.
    Date 2022-10-21
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://academic.oup.com/jhered/article/113/5/500/6657699
    Accessed 5/21/2025, 4:10:31 PM
    Rights https://creativecommons.org/licenses/by/4.0/
    Volume 113
    Pages 500-515
    Publication Journal of Heredity
    DOI 10.1093/jhered/esac038
    Issue 5
    ISSN 0022-1503, 1465-7333
    Date Added 5/21/2025, 4:10:31 PM
    Modified 5/21/2025, 4:10:31 PM

    Attachments

    • Etherington et al. - 2022 - Extensive genome introgression between domestic ferret and European polecat during population recove.pdf
  • Feral ferrets (Mustela furo) as hosts and sentinels of tuberculosis in New Zealand

    Item Type Journal Article
    Author A. E. Byrom
    Author P. Caley
    Author B. M. Paterson
    Author G. Nugent
    Abstract The control and eventual eradication of bovine tuberculosis (TB) poses major challenges in New Zealand, given the variety of wildlife species susceptible to TB, many of which are capable of onwards transmission of Mycobacterium bovis infection. Here we discuss the role of feral ferrets (Mustela furo), focussing on potential transmission or risk pathways that have implications for management of TB. Firstly inter-specific transmission to ferrets. Ferrets scavenge potentially infected wildlife, including other ferrets, thus prevalence of TB can be amplified through ferrets feeding on tuberculous carcasses, particularly brushtail possums (Trichosurus vulpecula). Secondly intra-specific transmission between ferrets. The rate of ferret-ferret transmission depends on population density, and in some places ferret densities exceed the estimated threshold for disease persistence. TB can therefore potentially be maintained independently of other sources of infection. Thirdly transmission from ferrets to other wildlife. These include the main wildlife maintenance host, brushtail possums, that will occasionally scavenge potentially tuberculous ferret carcasses. Fourthly transmission from ferrets to livestock. This is considered to occur occasionally, but the actual rate of transmission has never been measured. Fifthly geographical spread. M. bovis-infected ferrets can travel large distances and cause new outbreaks of TB at locations previously free of TB, which may have caused an expansion of TB-endemic areas.Ferrets play a complex role in the TB cycle in New Zealand; they are capable of contracting, amplifying and transmitting M. bovis infection, sometimes resulting in ferret populations with a high prevalence of TB. However, ferret population densities are usually too low to sustain infection independently, and transmission to other wildlife or livestock appears a rarer event than with possums. Nevertheless, management of ferrets remains a key part of the National Pest Management Strategy for TB. Control is prudent where M. bovis-infected ferret populations exist in high numbers, to reduce the onward transmission risk of any self-sustained infection to livestock. When ferret numbers are well below the theoretical disease maintenance threshold, ferret control is still sometimes warranted because of the animals’ ability to acquire infection when young and, through dispersal, transport it outside TB-endemic areas. Ferrets can also be used as disease sentinels for TB, especially in areas where alternative sentinel species are rare or expensive to survey, and when sampling of possums is not cost-effective.
    Date March 25, 2015
    Language en
    Library Catalog Taylor and Francis+NEJM
    URL https://doi.org/10.1080/00480169.2014.981314
    Accessed 4/29/2019, 10:51:02 AM
    Extra PMID: 25495945
    Volume 63
    Pages 42-53
    Publication New Zealand Veterinary Journal
    DOI 10.1080/00480169.2014.981314
    Issue sup1
    ISSN 0048-0169
    Date Added 4/29/2019, 10:51:02 AM
    Modified 5/21/2025, 4:52:12 PM

    Tags:

    • wildlife
    • pathogen
    • New Zealand
    • Mycobacterium bovis
    • Mustela furo
    • disease sentinel
    • Disease surveillance
    • Erratum
    • ferret
    • tuberculosis

    Attachments

    • Byrom et al. - 2015 - Feral ferrets (Mustela furo) as hosts and sentinel.pdf
    • Snapshot
  • Ferret behavior

    Item Type Journal Article
    Author Peter G. Fisher
    Date 2006
    Library Catalog PubMed Central
    URL https://www.ncbi.nlm.nih.gov/pmc/articles/PMC7158301/
    Accessed 2/20/2025, 5:59:00 PM
    Extra PMID: null PMCID: PMC7158301
    Pages 163-205
    Publication Exotic Pet Behavior
    DOI 10.1016/B978-1-4160-0009-9.50011-6
    Journal Abbr Exotic Pet Behavior
    Date Added 2/20/2025, 5:59:00 PM
    Modified 2/20/2025, 5:59:30 PM

    Attachments

    • PubMed Central Full Text PDF
    • PubMed Central Link
  • Ferrets: a selective overview of issues and options

    Item Type Report
    Author Kenneth W. Umbach
    Date May 30, 1997
    URL https://www.legalizeferrets.org/wp-content/uploads/2016/03/CA-Research-Library-Article.pdf
    Accessed 7/19/2023, 10:39:07 AM
    Place Sacramento, CA
    Report Number 3
    Report Type CRB Note
    Institution California Research Bureau
    Date Added 7/19/2023, 10:39:39 AM
    Modified 10/22/2025, 11:29:37 AM

    Attachments

    • Umbach - 1997 - Ferrets a selective overview of issues and options.pdf
  • Founder events, isolation, and inbreeding: Intercontinental genetic structure of the domestic ferret

    Item Type Journal Article
    Author Kyle D. Gustafson
    Author Michelle G. Hawkins
    Author Tracy L. Drazenovich
    Author Robert Church
    Author Susan A. Brown
    Author Holly B. Ernest
    Abstract Domestication and breeding for human-desired morphological traits can reduce population genetic diversity via founder events and artificial selection, resulting in inbreeding depression and genetic disorders. The ferret (Mustela putorius furo) was domesticated from European polecats (M. putorius), transported to multiple continents, and has been artificially selected for several traits. The ferret is now a common pet, a laboratory model organism, and feral ferrets can impact native biodiversity. We hypothesized global ferret trade resulted in distinct international genetic clusters and that ferrets transported to other continents would have lower genetic diversity than ferrets from Europe because of extreme founder events and no hybridization with wild polecats or genetically diverse ferrets. To assess these hypotheses, we genotyped 765 ferrets at 31 microsatellites from 11 countries among the continents of North America, Europe, and Australia and estimated population structure and genetic diversity. Fifteen M. putorius were genotyped for comparison. Our study indicated ferrets exhibit geographically distinct clusters and highlights the low genetic variation in certain countries. Australian and North American clusters have the lowest genetic diversities and highest inbreeding metrics whereas the United Kingdom (UK) cluster exhibited intermediate genetic diversity. Non-UK European ferrets had high genetic diversity, possibly a result of introgression with wild polecats. Notably, Hungarian ferrets had the highest genetic diversity and Hungary is the only country sampled with two wild polecat species. Our research has broad social, economic, and biomedical importance. Ferret owners and veterinarians should be made aware of potential inbreeding depression. Breeders in North America and Australia would benefit by incorporating genetically diverse ferrets from mainland Europe. Laboratories using ferrets as biomedical organisms should consider diversifying their genetic stock and incorporating genetic information into bioassays. These results also have forensic applications for conserving the genetics of wild polecat species and for identifying and managing sources of feral ferrets causing ecosystem damage.
    Date 2018
    Language en
    Short Title Founder events, isolation, and inbreeding
    Library Catalog Wiley Online Library
    URL https://onlinelibrary.wiley.com/doi/abs/10.1111/eva.12565
    Accessed 2/20/2025, 6:00:04 PM
    Rights © 2017 The Authors. Evolutionary Applications published by John Wiley & Sons Ltd
    Extra _eprint: https://onlinelibrary.wiley.com/doi/pdf/10.1111/eva.12565
    Volume 11
    Pages 694-704
    Publication Evolutionary Applications
    DOI 10.1111/eva.12565
    Issue 5
    ISSN 1752-4571
    Date Added 2/20/2025, 6:00:04 PM
    Modified 6/14/2025, 9:34:34 AM

    Tags:

    • Europe
    • Australia
    • North America
    • New Zealand
    • artificial selection
    • Mustela putorius
    • Mustela putorius furo

    Attachments

    • Full Text PDF
    • Snapshot
  • Genetic divergence without spatial isolation in polecat Mustela putorius populations

    Item Type Journal Article
    Author T. Lodé
    Date 2001
    Language en
    Library Catalog Google Scholar
    URL https://academic.oup.com/jeb/article-abstract/14/2/228/7322855
    Accessed 10/22/2025, 7:54:17 AM
    Extra Publisher: Blackwell Publishing Ltd Oxford, UK
    Volume 14
    Pages 228–236
    Publication Journal of Evolutionary Biology
    DOI 10.1046/j.1420-9101.2001.00275.x
    Issue 2
    ISSN 1420-9101
    Date Added 10/22/2025, 7:54:17 AM
    Modified 10/22/2025, 8:01:10 AM

    Tags:

    • genetic divergence
    • endogamy
    • habitat preferences
    • Mustela
    • polymorphism
    • sympatric differentiation

    Attachments

    • Full Text PDF
    • Snapshot
  • Genetic diversity and fitness in black-footed ferrets before and during a bottleneck

    Item Type Journal Article
    Author S. M. Wisely
    Date 2002-7-1
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://academic.oup.com/jhered/article-lookup/doi/10.1093/jhered/93.4.231
    Accessed 5/21/2025, 4:09:56 PM
    Volume 93
    Pages 231-237
    Publication Journal of Heredity
    DOI 10.1093/jhered/93.4.231
    Issue 4
    ISSN 14718505
    Date Added 5/21/2025, 4:09:58 PM
    Modified 5/21/2025, 4:13:16 PM

    Attachments

    • Wisely - 2002 - Genetic diversity and fitness in black-footed ferrets before and during a bottleneck.pdf
  • Genetic evaluation of a reintroduced population of black-footed ferrets (Mustela nigripes)

    Item Type Journal Article
    Author Cynthia M. Cain
    Author Travis M. Livieri
    Author Bradley J. Swanson
    Date 2011-08-16
    Language en
    Library Catalog CrossRef
    URL http://www.bioone.org/doi/abs/10.1644/10-MAMM-S-104.1
    Accessed 10/6/2014, 1:19:31 PM
    Volume 92
    Pages 751-759
    Publication Journal of Mammalogy
    DOI 10.1644/10-MAMM-S-104.1
    Issue 4
    ISSN 0022-2372, 1545-1542
    Date Added 10/6/2014, 1:19:31 PM
    Modified 5/21/2025, 4:53:01 PM

    Attachments

    • Cain et al. - 2011 - Genetic evaluation of a reintroduced population of.pdf
  • Genotypic and phenotypic consequences of reintroduction history in the black-footed ferret (Mustela nigripes)

    Item Type Journal Article
    Author Samantha M. Wisely
    Author Rachel M. Santymire
    Author Travis M. Livieri
    Author Sara A. Mueting
    Author JoGayle Howard
    Date 4/2008
    Language en
    Library Catalog CrossRef
    URL http://link.springer.com/10.1007/s10592-007-9351-x
    Accessed 10/6/2014, 1:16:47 PM
    Volume 9
    Pages 389-399
    Publication Conservation Genetics
    DOI 10.1007/s10592-007-9351-x
    Issue 2
    ISSN 1566-0621, 1572-9737
    Date Added 10/6/2014, 1:16:47 PM
    Modified 5/21/2025, 8:35:50 PM

    Attachments

    • Wisely et al. - 2008 - Genotypic and phenotypic consequences of reintrodu.pdf
  • Home-range size and spatial organization of black-footed ferrets Mustela nigripes in South Dakota, USA

    Item Type Journal Article
    Author David S. Jachowski
    Author Joshua J. Millspaugh
    Author Dean E. Biggins
    Author Travis M. Livieri
    Author Marc R. Matchett
    Date 03/2010
    Language en
    Library Catalog CrossRef
    URL http://www.bioone.org/doi/abs/10.2981/09-034
    Accessed 7/29/2014, 10:39:48 AM
    Volume 16
    Pages 66-76
    Publication Wildlife Biology
    DOI 10.2981/09-034
    Issue 1
    ISSN 0909-6396
    Date Added 7/29/2014, 10:39:48 AM
    Modified 5/21/2025, 8:36:07 PM

    Attachments

    • Jachowski et al. - 2010 - Home-range size and spatial organization of black-.pdf
  • Hybridization and the phylogenetic relationship between polecats and domestic ferrets in Britain

    Item Type Journal Article
    Author A Davison
    Author J.D.S Birks
    Author H.I Griffiths
    Author A.C Kitchener
    Author D Biggins
    Author R.K Butlin
    Abstract Ferrets (Mustela furo) were domesticated from polecats (M. putorius, M. eversmannii) over 2000 years ago. Following their introduction to Britain, they escaped and hybridized with native European polecats (M. putorius). Native polecats declined to the point of near extinction prior to World War I, but have recently begun to expand from a Welsh refugium. Concern has arisen as to the extent of polecat/ferret introgression, and in particular, whether the expanding population is of mainly hybrid origin. Therefore, mitochondrial DNA sequencing was used to investigate polecat genetic diversity in Britain. Two geographically distinct lineages were found, where one may be ancestral to the British polecat, and the other to the domestic ferret. The ancestral distribution of each lineage, or assortative mating is su•cient to explain the observed pattern. A further comparison between the distribution of the polecat phenotype and mitochondrial haplotype implies that the current population expansion may be mediated by dispersing male polecats hybridizing with female feral ferrets. However, the wild source of the ferret remains obscure. Relatively recent speciation from European mink (M. lutreola) and black-footed ferrets (M. nigripes), and/or the e€ects of hybridization result in an unresolved molecular phylogeny. # 1998 Elsevier Science Ltd. All rights reserved.
    Date 02/1999
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://linkinghub.elsevier.com/retrieve/pii/S0006320798000676
    Accessed 2/20/2025, 6:06:52 PM
    Rights https://www.elsevier.com/tdm/userlicense/1.0/
    Volume 87
    Pages 155-161
    Publication Biological Conservation
    DOI 10.1016/S0006-3207(98)00067-6
    Issue 2
    Journal Abbr Biological Conservation
    ISSN 00063207
    Date Added 2/20/2025, 6:06:52 PM
    Modified 6/14/2025, 9:34:25 AM

    Attachments

    • Davison et al. - 1999 - Hybridization and the phylogenetic relationship between polecats and domestic ferrets in Britain.pdf
  • Identifying behavioural traits and underlying personality dimensions in domestic ferrets (Mustela putorius furo)

    Item Type Journal Article
    Author Sarah Talbot
    Author Rafael Freire
    Author Skye Wassens
    Abstract The aim of this study was to examine the personality structure of domestic ferrets (Mustela putorius furo) by using owner-based reporting of personality traits. A total of 743 ferret owners participated in an online questionnaire, with a total of 1029 ferrets being assessed. Respondents rated 67 adjectives based on their ferret(s) behavioural traits and personality. Principal component analysis (PCA) of these trait ratings identified four underlying personality dimensions, which accounted for 47.1% of the total variance. These were labelled according to the traits that they encompass: Extraversion, Sociability, Attentiveness, and Neuroticism. Details about ferret sex, de-sexing status, age, and coat colour were also sought, and General Linear Mixed Models were used to test the main effects of these characteristics on the personality dimensions. It was found that sex (p < 0.01) and age (p < 0.001) significantly influenced certain personality components, whereas de-sexing did not. Sociability, Attentiveness, and Neuroticism were found to differ based on sex, whereby males were rated as more sociable than females, but females were rated higher on the Attentiveness and Neuroticism subscales. Finally, Extraversion was found to generally decrease with age. We can use the findings of this study to make cross-species comparisons and further inform the discussion regarding the adaptive relevance of animal personality. Identifying differences in personality types can improve welfare by using this information to satisfy the different needs of individuals.
    Date 2021/8
    Language en
    Library Catalog www.mdpi.com
    URL https://www.mdpi.com/2076-2615/11/8/2173
    Accessed 2/20/2025, 6:05:11 PM
    Rights http://creativecommons.org/licenses/by/3.0/
    Extra Number: 8 Publisher: Multidisciplinary Digital Publishing Institute
    Volume 11
    Pages 2173
    Publication Animals
    DOI 10.3390/ani11082173
    Issue 8
    ISSN 2076-2615
    Date Added 2/20/2025, 6:05:11 PM
    Modified 10/21/2025, 4:40:28 PM

    Tags:

    • behaviour
    • welfare
    • animal personality
    • domestic ferrets

    Attachments

    • Full Text PDF
  • Impacts of domesticated ferrets upon wildlife, agriculture, and human health in the USA, compiled from state agency surveys and literature review, with special emphasis upon California

    Item Type Journal Article
    Author Gary Orval Graening
    Date 2022-7-1
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://wildlife.ca.gov/Publications/Journal/Issues/volume-108-issue-2-impacts-of-domesticated-ferrets-upon-wildlife-agriculture-and-human-health-in-the-usa-compiled-from-state-agency-surveys-and-literature-
    Accessed 11/9/2022, 8:49:19 AM
    Volume 108
    Publication California Fish and Wildlife Journal
    DOI 10.51492/cfwj.108.13
    Issue 2
    ISSN 2689-4203, 2689-419X
    Date Added 11/9/2022, 8:49:22 AM
    Modified 10/22/2025, 11:30:35 AM

    Attachments

    • Graening - 2022 - Impacts of domesticated ferrets upon wildlife, agr.pdf
    • Snapshot
  • Intraspecific and seasonal differences in the diet of feral ferrets (Mustela furo) in a pastoral habitat, East Otago, New Zealand

    Item Type Journal Article
    Author J R Ragg
    Date 1998
    Language en
    Library Catalog Zotero
    Volume 22
    Pages 113-119
    Publication New Zealand Journal of Ecology
    Issue 2
    Date Added 4/29/2019, 10:52:18 AM
    Modified 5/21/2025, 8:34:16 PM

    Attachments

    • Ragg - 1998 - Intraspecific and seasonal differences in the diet.pdf
  • Investigating infectious disease threats to the recovery of the European polecat in Britain

    Item Type Journal Article
    Author Kari-Anne Heald
    Author Caroline Millins
    Author Andrew C. Kitchener
    Author Ashley C. Banyard
    Author Georg Hantke
    Author Katherine A. Sainsbury
    Author Michael McDonald
    Author Anna Meredith
    Date 08/2020
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://link.springer.com/10.1007/s42991-020-00046-6
    Accessed 10/22/2025, 7:54:09 AM
    Volume 100
    Pages 439-444
    Publication Mammalian Biology
    DOI 10.1007/s42991-020-00046-6
    Issue 4
    Journal Abbr Mamm Biol
    ISSN 1616-5047, 1618-1476
    Date Added 10/22/2025, 7:54:17 AM
    Modified 10/22/2025, 7:54:17 AM

    Attachments

    • Available Version (via Google Scholar)
  • Isolation and Characterization of Microsatellite Markers in the Domestic Ferret (Mustela putorius furo)

    Item Type Journal Article
    Author Holly Ernest
    Author Tracy Drazenovich
    Author Lisa Dalbeck
    Author Michelle Hawkins
    Abstract The domestic ferret (Mustela putorius furo) is an important model organism for the study of avian influenza and other diseases of humans and animals, as well as a popular pet animal. In order to evaluate genetic diversity and study disease relationships in ferrets, 22 nuclear microsatellite loci (17 dinucleotide and 5 tetranucleotide) were developed from ferret genomic libraries and organized into seven multiplex sets. Polymorphism was preliminarily assessed in one population in Australia and one in the USA, sampled with 25 individuals each. The loci displayed allelic diversity ranging from 1 to 5 alleles, and expected and observed heterozygosities ranging from 0.04 to 0.65 and 0.04 to 0.76, respectively. Additionally, the loci amplified products in 15 samples from the wild ancestor, European polecat (Mustela putorius) and domestic ferret-polecat hybrids. In polecat/hybrid samples, allelic diversity ranged from 3 to 8 alleles, and expected and observed heterozygosities ranged from 0.13 to 0.81 and 0.13 to 0.80 respectively. These markers will be useful for molecular assessments of genetic diversity and applications to evolution, ecology, and health in domestic ferrets and wild polecats.
    Date 2012-12-05
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://www.mdpi.com/1422-0067/13/12/16592
    Accessed 5/21/2025, 4:10:45 PM
    Rights https://creativecommons.org/licenses/by/3.0/
    Volume 13
    Pages 16592-16597
    Publication International Journal of Molecular Sciences
    DOI 10.3390/ijms131216592
    Issue 12
    Journal Abbr IJMS
    ISSN 1422-0067
    Date Added 5/21/2025, 4:10:45 PM
    Modified 5/21/2025, 4:10:45 PM

    Attachments

    • Ernest et al. - 2012 - Isolation and Characterization of Microsatellite Markers in the Domestic Ferret (Mustela putorius fu.pdf
  • Man’s underground best friend: domestic ferrets, unlike the wild forms, show evidence of dog-like social-cognitive skills

    Item Type Journal Article
    Author Anna Hernádi
    Author Anna Kis
    Author Borbála Turcsán
    Author József Topál
    Abstract Recent research has shown that dogs’ possess surprisingly sophisticated human-like social communication skills compared to wolves or chimpanzees. The effects of domestication on the emergence of socio-cognitive skills, however, are still highly debated. One way to investigate this is to compare socialized individuals from closely related domestic and wild species. In the present study we tested domestic ferrets (Mustela furo) and compared their performance to a group of wild Mustela hybrids and to domestic dogs (Canis familiaris). We found that, in contrast to wild Mustela hybrids, both domestic ferrets and dogs tolerated eye-contact for a longer time when facing their owners versus the experimenter and they showed a preference in a two-way choice task towards their owners. Furthermore, domestic ferrets, unlike the wild hybrids, were able to follow human directional gestures (sustained touching; momentary pointing) and could reach the success rate of dogs. Our study provides the first evidence that domestic ferrets, in a certain sense, are more dog-like than their wild counterparts. These findings support the hypothesis that domestic species may share basic socio-cognitive skills that enable them to engage in effectively orchestrated social interactions with humans.
    Date Aug 15, 2012
    Language en
    Short Title Man’s underground best friend
    Library Catalog PLoS Journals
    URL https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0043267
    Accessed 10/4/2019, 3:38:12 PM
    Volume 7
    Pages e43267
    Publication Plos One
    DOI 10.1371/journal.pone.0043267
    Issue 8
    Journal Abbr PLOS ONE
    ISSN 1932-6203
    Date Added 10/4/2019, 3:38:12 PM
    Modified 10/22/2025, 11:30:30 AM

    Tags:

    • Animal behavior
    • Animal performance
    • Animal sociality
    • Dogs
    • Domestic animals
    • Ferrets
    • Pets and companion animals
    • Social communication
    • Wolves

    Attachments

    • Hernádi et al. - 2012 - Man’s underground best friend domestic ferrets, u.pdf
    • Snapshot
  • Microhabitat selection by feral ferrets (Mustela furo) in a pastoral habitat, East Otago, New Zealand

    Item Type Journal Article
    Author J. R. Ragg
    Author H. Moller
    Abstract [The spatial distribution of feral ferret (Mustela furo) activity and denning were studied using ink-print tracking tunnels and radio-tracking within pastoral farmland containing a mosaic of grazed (developed and semi-developed) and ungrazed pasture, scrub, tree plantation and scrubby fence lines at Palmerston, East Otago, South Island, New Zealand. Ferrets concentrated their activity in grazed areas but within these areas they were found more often where herbs, scrub and woody cover were present, and where there was an ecotone between pasture and vegetation cover. Ferrets were more likely to be present close to fence lines. When denning, ferrets selected areas with cover of all types and avoided open pasture areas. Ferrets particularly favoured man-made structures (woolsheds, haybarns, sheds etc.) for dens. The implications of these microhabitat selections by ferrets to wildlife conservation in New Zealand are discussed. Ferrets are a possible vector of bovine tuberculosis (Mycobacterium bovis). Ferrets may transmit tuberculosis to stock via contaminated food or latrines deposited outside den sites (68% of dens were accessible to stock). Concentration of ferret movements along pasture ecotones may be a factor facilitating tuberculosis transmission between possums (Trichosurus vulpecula) and ferrets, and between ferrets and stock.]
    Date 2000
    Archive JSTOR
    Library Catalog JSTOR
    URL https://www.jstor.org/stable/24054649
    Accessed 4/29/2019, 10:47:41 AM
    Volume 24
    Pages 39-46
    Publication New Zealand Journal of Ecology
    Issue 1
    ISSN 0110-6465
    Date Added 4/29/2019, 10:47:41 AM
    Modified 5/21/2025, 8:44:57 PM

    Attachments

    • Ragg and Moller - 2000 - Microhabitat selection by feral ferrets (Mustela f.pdf
  • Mortality and reproductive patterns of wild European polecats Mustela putorius in Denmark

    Item Type Journal Article
    Author Lise V. Kristiansen
    Author Peter Sunde
    Author Gösta Nachman
    Author Aksel B. Madsen
    Date 12/2007
    Language en
    Library Catalog DOI.org (Crossref)
    URL http://link.springer.com/10.1007/BF03194235
    Accessed 10/22/2025, 7:55:28 AM
    Rights http://www.springer.com/tdm
    Volume 52
    Pages 371-378
    Publication Acta Theriologica
    DOI 10.1007/BF03194235
    Issue 4
    Journal Abbr Acta Theriol
    ISSN 0001-7051, 2190-3743
    Date Added 10/22/2025, 7:55:28 AM
    Modified 10/22/2025, 7:55:58 AM

    Attachments

    • PDF
  • Mortality rates of feral ferrets (Mustela furo) in New Zealand

    Item Type Journal Article
    Author Peter Caley
    Author L. M. McElrea
    Author Jim Hone
    Abstract Life-table data from feral ferret populations in New Zealand were analysed to estimate their mortality rates, and to test for any additive effect of Mycobacterium bovis infection on observed mortality rates. The observed instantaneous mortality rate was best estimated by modelling mortality as a 2-phase step model with different rates for juveniles (μ1 = 1.45 year–1, 95% C.I. 1.2–1.7 year–1) and adults (μ2 = 0.55 year–1, 95% C.I. 0.4–0.9 year–1). This corresponds to a survival probability of 0.25 during the first year of life, rising to 0.55 year–1 thereafter, and a life expectancy of 0.95 years. At a population level, no additional mortality due to M. bovis infection was observed, suggesting either that the rate of disease-induced mortality was negligible, or that it was compensatory with natural mortality.
    Date 2002
    Language en
    Library Catalog www.publish.csiro.au
    URL http://www.publish.csiro.au/WR/WR02004
    Accessed 4/29/2019, 10:48:56 AM
    Volume 29
    Pages 323-328
    Publication Wildlife Research
    DOI 10.1071/wr02004
    Issue 4
    Journal Abbr Wildl. Res.
    ISSN 1448-5494
    Date Added 4/29/2019, 10:48:56 AM
    Modified 5/21/2025, 8:44:48 PM

    Attachments

    • Caley et al. - 2002 - Mortality rates of feral ferrets (Mustela furo) in New Zealand.pdf
    • Snapshot
  • Movements and habitat use of feral house cats Felis catus, stoats Mustela erminea and ferrets Mustela furo, in grassland surrounding Yellow-eyed penguin Megadyptes antipodes breeding areas in spring

    Item Type Journal Article
    Author N. Alterio
    Author H. Moller
    Author H. Ratz
    Date 02/1998
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://linkinghub.elsevier.com/retrieve/pii/S0006320797000529
    Accessed 4/29/2019, 10:52:04 AM
    Volume 83
    Pages 187-194
    Publication Biological Conservation
    DOI 10.1016/S0006-3207(97)00052-9
    Issue 2
    Journal Abbr Biological Conservation
    ISSN 00063207
    Date Added 4/29/2019, 10:52:04 AM
    Modified 5/21/2025, 8:44:36 PM

    Attachments

    • Alterio et al. - 1998 - Movements and habitat use of feral house cats Feli.pdf
  • Notes on the genetic behaviour of certain characters in the polecat, ferret, and in polecat-ferret hybrids

    Item Type Journal Article
    Author Frances Pitt
    Date 9/1921
    Language en
    Library Catalog DOI.org (Crossref)
    URL http://link.springer.com/10.1007/BF02983044
    Accessed 5/21/2025, 4:10:47 PM
    Rights http://www.springer.com/tdm
    Volume 11
    Pages 95-115
    Publication Journal of Genetics
    DOI 10.1007/BF02983044
    Issue 2
    Journal Abbr Journ. of Gen.
    ISSN 0022-1333
    Date Added 5/21/2025, 4:10:47 PM
    Modified 5/21/2025, 4:10:47 PM

    Attachments

    • Pitt - 1921 - Notes on the genetic behaviour of certain characters in the polecat, ferret, and in polecat-ferret h.pdf
  • On the lack of a universal pattern associated with mammalian domestication: differences in skull growth trajectories across phylogeny

    Item Type Journal Article
    Author Marcelo R. Sánchez-Villagra
    Author Valentina Segura
    Author Madeleine Geiger
    Author Laura Heck
    Author Kristof Veitschegger
    Author David Flores
    Abstract As shown in a taxonomically broad study, domestication modifies postnatal growth. Skull shape across 1128 individuals was characterized by 14 linear measurements, comparing 13 pairs of wild versus domesticated forms. Among wild forms, the boar, the rabbit and the wolf have the highest proportion of allometric growth, explaining in part the great morphological diversity of the domesticated forms of these species. Wild forms exhibit more isometric growth than their domesticated counterparts. Multivariate comparisons show that dogs and llamas exhibit the greatest amount of differences in trajectories with their wild counterparts. The least amount is recorded in the pig–boar, and camel and horse pairs. Bivariate analyses reveal that most domesticated forms have growth trajectories different from their respective wild counterparts with regard to the slopes. In pigs and camels slopes are shared and intercepts are different. There is a trajectory extension in most domesticated herbivores and the contrary pattern in carnivorous forms. However, there is no single, universal and global pattern of paedomorphosis or any other kind of heterochrony behind the morphological diversification that accompanies domestication.
    Date 2017-10-25
    Short Title On the lack of a universal pattern associated with mammalian domestication
    Library Catalog royalsocietypublishing.org (Atypon)
    URL https://royalsocietypublishing.org/doi/10.1098/rsos.170876
    Accessed 7/14/2025, 4:15:03 PM
    Extra Publisher: Royal Society
    Volume 4
    Pages 170876
    Publication Royal Society Open Science
    DOI 10.1098/rsos.170876
    Issue 10
    Date Added 7/14/2025, 4:15:03 PM
    Modified 7/14/2025, 4:15:03 PM

    Tags:

    • modularity
    • dog
    • development
    • ontogeny
    • cat
    • horse

    Attachments

    • Full Text PDF
  • Pet European ferrets: a hazard to public health, small livestock and wildlife

    Item Type Report
    Author Denny G. Constantine
    Author Kenneth W. Kizer
    Date Dec 1988
    Library Catalog Google Scholar
    Extra Publisher: BMJ Publishing Group
    Place Sacramento, CA
    Pages 71
    Institution California Department of Health Services
    Date Added 7/14/2023, 12:01:57 PM
    Modified 5/22/2025, 8:01:59 PM

    Notes:

    • wl cal

      PET EUROPEAN FERRETS:
      A HAZARD TO PUBLIC HEALTH,

      SMALL LIVESTOCK AND WILDLIFE

      PET EUROPEAN FERRETS: A HAZARD TO

      PUBLIC HEALTH, SMALL LIVESTOCK AND WILDLIFE

      Denny G. Constantine, D.V.M., M.P.H.
      Public Health Veterinarian
      Veterinary Public Health Unit

      and

      Kenneth W. Kizer, M.D., M.P.H.
      Director

      California Department of Health Services
      714 P Street
      Sacramento, California 95814

      December 1988
      EXECUTIVE SUMMARY

      European ferrets are cute and playful, and they are becoming
      increasingly popular as pets. However, their growing popularity is viewed
      with alarm by numerous public health entities and animal care organizations.

      Ferret play frequently assumes the form of mock attacks, which may
      result in bites to humans. Serious bites may occur, especially if the
      animal is surprised or angered. Adults are able to quickly terminate such
      encounters, and thereby limit injury. However, infants, who often seem to
      be perceived by ferrets as prey, may suffer severe injury as a result of
      ferret attacks. Indeed, ferrets sometimes unleash frenzied, rapid-fire bite
      and slash attacks on infants, usually on their heads and throats, and
      sometimes inflict hundreds of bites. The animals have been reported to then
      drink the victim's blood and eat the shredded tissues.

      In order to better define the nature and extent of ferret attacks, and
      in response to requests for information from other state agencies, the
      California Department of Health Services solicited reports about ferret
      attacks in early 1986. During the subsequent 2 years, information was
      obtained on 452 ferret attacks spanning the 10-year period 1978 through
      1987. 425 attacks on people were reported from California, Oregon, and
      Arizona. One hundred of these attacks were from California, where it is
      illegal to keep ferrets as pets. Also reported from a total of 18 states
      were 63 unprovoked attacks on infants and small children. Several of these
      were near fatal attacks. One additional case, a fatal attack, was reported

      from London, England.
      Data from California indicate that the majority of attacks were
      inflicted by pet ferrets belonging to households other than the victim's.

      Twenty-eight percent of infants required plastic and reconstructive
      surgery; 22 percent of victims required rabies prophylaxis; and 4 percent of
      victims were known to have been exposed to rabid ferrets.

      Ferrets have a propensity for escaping from their principle residence,
      and escaped ferrets are known to boldly approach wildlife. These ferrets

      may develop rabies after returning home. Twelve such cases have been
      reported in the United States.

      Ferrets develop feral populations and are especially destructive of
      poultry and small wild animals such as rabbits. As a result of this well
      recognized problem, the keeping of ferrets as pets was outlawed in
      California in 1935. However, data gathered from our survey indicate that

      greater surveillance and enforcement efforts in this regard may be needed.

      -ii-
      II.

      IIl.

      IV.

      TABLE OF CONTENTS

      Executive Summary ........ cee eccccceeee ee er
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      B. Attacks on Persons in California ........ cc cece eee enesees
      C. Attacks on Persons in California, Oregon,
      and Arizona .... cece eee e cence neces PERSE WERT eee di Soba dus
      D. Ferret Rabies in the United States .......... cece ee eens
      E. Feral Ferrets in California TTTTERT CLE

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      Rabies in Pet Ferrets ....... ia fa (910 a! ow ie We te 9 6) al mi fate 19 91 8h gi wn He
      Attacks According to Age of Victim ........... TTT eT
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      Liability of Ferret Ownership .........cccceecescccecsence
      Feral Ferret Populations Can Devastate Small

      Livestock and Native Vertebrates and Contribute

      to the Wildlife Rabies Problem .............. scene eens cece

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      Acknowledgments ........ eee re wese db bb beoan ded FATTTT ES RETRO
      References .. cece cere c cece rece reece new n ere eee e ree eters trees

      Appendix I. California State Law Relevant to Ferrets........ aie
      I. INTRODUCTION

      The European ferret, Mustela putorius, is cute and playful, and it has
      become an increasingly popular pet in recent years. However, reports of
      ferret attacks on people, especially infants, also have increased in recent
      years. This report briefly describes the history of the European ferret and
      reports the findings of a survey of ferret attacks conducted by the
      California Department of Health Services (CDHS).

      Developed from the European polecat, Mustela putorius (Volobuev, et al,
      1974), European ferrets resemble weasels, except for being larger and
      stockier. Ferrets--along with the wolverine, weasel, marten, and mink--are
      members of the family Mustelidae and subfamily Mustelinae. They measure 17
      to 22 inches in length and typically weigh between 2 and 6 pounds. Most
      ferrets are albino or "sable" colored (yellow-buff undercoat overlaid with
      blackish guard hairs) with a blackish mask, limbs, and tail, although other
      colors exist. The European ferret should not be confused with the similar
      but biologically distinct American black-footed ferret, Mustela nigripes, an
      endangered species of the Great Plains.

      Since at least 63 B.C., ferrets have been used by man to drive rabbits
      and rats from their holes (Owen, 1969). They are preferred for this because
      of the fierce and relentless nature of their attacks (Everitt, 1897). Only
      recently have ferrets become popular as household pets not expected to
      engage in the "ferreting" activities that have characterized the animal's
      long relationship with man. And while ferrets may be cute and playful, they
      are frequent biters, sometimes inflicting bites with machine gun rapidity

      and occasionally tenaciously refusing to let go of their victim. These
      traits have been recently reported in numerous accounts of ferret attacks on
      human infants, although the propensity of ferrets "to attack and kill
      children in the cradle" is longstanding (Fennell, 1841).

      Our first acquaintance with problems concerning pet ferrets occurred in
      early 1986, consequent to requests from two other departments of California
      State Government. The Department of Fish and Game (DFG) and the
      Department of Food and Agriculture (DFA) asked DHS to define any public
      health concerns relevant to pet ferrets. DFG had restrictions against entry
      of pet ferrets into the State (except for an occasional castrated male, an
      exception since deleted), due to knowledge that escaped ferrets develop
      feral populations that have a tendency to ravage wildlife and small
      livestock (e.g., poultry and rabbits).

      Since 1986, California's ferret restrictions, along with those of other
      states, have been under attack by proponents of pet ferrets. Apparently,
      this is part of a national campaign to popularize and sell the animals
      (Anonymous, 1986). Ferret proponents claim that: (1) ferrets are domestic,
      rather than wild, and therefore, they should not be restricted; (2) ferrets
      bite people less often than dogs, so they are less hazardous; (3) pet
      ferrets are kept indoors, so there is no danger that they will get rabies
      from wildlife; and (4) ferrets will die if they escape and are not cared for
      by man.

      In an effort to respond to the DFG and DFA, we reviewed historical and
      recent literature on ferrets, and we solicited information on ferret attacks

      on people in order to better define the nature and extent of such events.
      In the process, we rediscovered and added to information that contradicts

      the claims of pet ferret proponents.

      II. METHODS

      Beginning in early 1986, we requested reports of ferret bites and
      attacks from all California counties and from two adjacent states. Requests
      for data from more distant states were also made concerning attacks on
      infants and rabies in ferrets. The medical literature was reviewed, as was
      literature regarding the habits of ferrets and their polecat progenitors and
      the existence of feral ferret populations.

      Early in the investigation it became apparent that some animal control
      personnel and laboratory workers needed help differentiating ferrets and
      weasels, whereupon a table somacicing differences was devised and provided
      to these persons.

      Other difficulties were experienced gathering and interpreting data.
      Ferret proponents made widely varying claims about the number of illegal pet
      ferrets existing in California (ranging from 100,000 to 500,000), and we
      lacked reliable figures on actual statewide numbers of either captive,
      stray, or feral ferrets. Owners of illegal ferrets generally were reluctant
      to report ferret attacks out of fear of prosecution or loss of their
      animals, and some bitten friends of owners were similarly reluctant. Most
      biting ferrets appeared to be strays or animals that had escaped or been
      released after biting their owners. Available reports, which were usually
      retrospective, generally lacked uniformity, and reflected varying degrees

      of effort in documenting details of the incident. Spectacular incidents,
      including those from rabid ferrets or involving infant maulings, seemed more
      likely to be reported.

      Rabies diagnostic tests were usually performed using the fluorescent
      rabies antibody test and mouse inoculation test (Johnson, 1979), although
      the monoclonal antibody technique (Wiktor and Koprowski, 1978) was used in
      some instances. When performed, the statistical significance of data
      associations were determined using the chi-square test, four-fold table

      method with Yates' correction.

      IIT. RESULTS

      Altogether, information was obtained on a total of 452 ferret attacks
      spanning the 10-year period 1978 through 1987. This included 64 unprovoked
      attacks on infants and young children and 388 attacks on older children and

      adults. The nature and details of these attacks are described below.

      A. Unprovoked Attacks on Infants

      Reports were received from 18 states and London, England, on
      unprovoked attacks on infants and young children (Tables 1 and 2).
      Thirty seven of these were from California (11), Arizona (17), and
      Oregon (9). All but one of the infants and small children were either
      reported to be, or appeared to be from bite report descriptions, three
      years of age or younger. One 6-year old is included among these cases
      because, upon being forced to release its hold, the ferret that seized
      the cheek of this sleeping girl ran and jumped into the crib of a
      subsequently rescued 16-month-old male sibling, each time biting the

      rescuing parent. The one fatal case was reported from London, although

      -~4-
      several of the other attacks were severe enough so as to nearly be
      fatal.

      The specific ages of the victims were reported for 50 of these
      youngsters, and ranged from 2 days to 3 years, with a median of 6
      months. Several of the most severely mauled infants were only a few
      weeks old.

      Gender was reported for 46 of the 64 infants; 24 (52 percent) were
      females.

      It appeared that as many as 58 (91 percent) of the victims were
      attacked while sleeping or lying down. Of the remaining six infants,
      one was bitten in the face by a rabid ferret from a pet shop. Another.
      was attacked on the wrist during a diaper change. A third was playing
      on the floor when bitten on its scalp. Two infants were outdoors when
      Stray ferrets attacked their feet. The last one was sitting on the
      toilet when the ferret jumped onto her face, inflicted multiple wounds,
      and then refused to open its jaws, requiring that it be pried loose.

      Wound sites were reported for 62 of the 63 injured infants. (The
      64th attack was blocked by a parent, so that the infant was not
      actually harmed.) Thirty (48 percent) infants suffered head wounds
      only; 11 (18 percent) received head or neck and limb wounds; 3 (5
      percent) were attacked on the neck only; and 18 (29 percent) were
      bitten just on their limbs or appendages. One infant was bitten on the
      end of his penis. Overall, 44 (71 percent) of the infants were bitten
      on the head or neck (plus limbs in 11 of these cases), whereas 18 (29

      percent) were bitten only on limbs or appendages. These data suggest
      that ferrets have a predilection for biting infants and small children
      on the head and/or neck.

      The sites of the head and neck wounds were identified in the
      reports (involving some overlap) as follows: head, 6; face, 23; cheek,
      7; eyelids, eyes, and conjunctiva, 5; lips or mouth, 4; scalp, 5;
      forehead, 2; nose, 1; ears, 3; neck, 3.

      Bites were usually described as consisting of multiple puncture
      wounds and lacerations, sometimes involving hundreds of bites per
      victim and leaving macerated tissues that resembled bloody ground beef.
      In 62 reports containing relevant information, 3 (5 percent) victims
      received a single bite or laceration, 14 (23 percent) were reported
      only as “bitten,” 44 (71 percent) received multiple bites and/or
      lacerations, and 1 (2 percent) had his ear eaten off. Ears were also
      bitten on 3 of the victims that received multiple facial bites, one of
      these having 40 percent of both ears eaten.

      The greatest reported numbers of multiple facial bites per victim
      were as follows: "hundreds," two; more than 200, one; 100, one; 80,
      one; 50, one; 40, one; and 20, one. The number of facial bites was not
      reported for one infant that exsanguinated and another that nearly bled
      to death. One 29-day-old infant lost her entire nose, and most of her
      eyelids, lips, and other facial tissues; her hands were also chewed. A
      2-day-old infant with some 80 puncture wounds on her head was reported
      by the mother as being "flipped over like a piece of meat" by a pet

      ferret that had its teeth fixed in the baby's scalp. In at least three
      instances, the jaws of the biting ferret had to be pried open to
      release the victim.

      Data on the characteristics of the ferrets attacking the infants
      are given in terms of numbers of persons attacked to underscore the
      assault perspective and to avoid confusion, because in two instances
      two persons were attacked by one ferret, and in two other instances
      one person was attacked by two ferrets. In one of the latter
      instances, ferret sex and age data were available and added to the
      appropriate columns of Table 2, resulting in a greater total number of
      ferrets (65) than assaulted infants (64). Seven (39 percent) of the
      ferrets whose ages were reported were less than one year old. The
      genders of the attacking ferrets were evenly divided between males and
      females, although this information was reported for only 14 animals.
      At least 8 of the 64 attacks were by unneutered animals.

      Data on the apparent ownership of the attacking animal were
      available for 48 of the 65 ferrets. These data indicated that 16 (33
      percent) attacks were from ferrets of the victim's household, whereas
      32 (67 percent) either had other known owners (25) or were strays (7).
      Of note, none of the 32 animals hesitated to approach people,
      suggesting that they were previously or currently pets. Six of the
      seven strays were friendly and had just been taken as household pets.
      Twenty-one of the 25 known owners were described as babysitters (6),
      visitors (7), hosts (5), and vendors (3) who had just supplied the
      ferrets. One (4 percent) of the 25 attacking animals that were tested

      for rabies was rabid.
      ae

      Data were reported on 100 ferret attacks in California (Table 3);
      this included 11 of the 64 infants reported above.

      The age of the victim was reported in 87 cases. Fourteen (16
      percent) were 3 years old or younger (3 of these were not unprovoked
      attacks), 9 (10 percent) were >3 to 10 years old, 13 (15 percent) were
      >10 to 20 years old, and 51 (59 percent) were older than 20 years. The
      oldest person was 70 years.

      The gender of the victims were approximately evenly divided, and
      similar in all age groups except for there being four males and ten
      females three years old or younger.

      Eleven (79 percent) of the 14 infants or children 3 years old or
      younger were attacked without provocation, 10 being in bed or asleep at
      the time of attack. A stray or feral ferret was found outdoors near a
      screaming three-year-old child whose feet and hand had been bitten.
      Three (19 percent) attacks occurred when handling or playing with stray
      or pet ferrets.

      According to the reported data, only 8 of 66 (12 percent) victims
      over 3 years of age were attacked without provocation. Illustrative of
      these cases was that of a seven-year-old girl who was bitten on the leg
      by a stray ferret that rushed from some bushes immediately after the
      girl had gotten out of an automobile. It had to be driven away, but it
      was later captured, killed, and tested for rabies.

      Seven adults were reported to have been attacked without
      provocation. The ankle of one was bitten, penetrating leather boots,

      by a feral or stray ferret. Another person's finger was bitten as he

      -8-
      tried to put trash into a dumpster, where the animal was hiding.
      Another man was bitten on his foot as he stepped onto his porch. A
      woman was bitten on the ankle by a stray ferret hiding in her garage.
      The feet and toes of two adults were bitten by pet ferrets; another
      adult was bitten several times in the Achilles' tendon by a stray
      ferret that had scratched on his front door. Two other feral or stray
      ferrets climbed onto persons and subsequently bit them. It was
      sometimes not clear from the reports whether the animals were friendly
      Or aggressive.

      The remaining 58 (88 percent) victims over 3 years of age, and 6
      of the 13 persons of unreported age, facilitated the attacks by
      handling, feeding, or in some way interacting with or provoking the
      animal. At least ten animal control officers were bitten trying to
      manipulate or care for ferrets.

      The following "victim-facilitated" or provoked cases are

      noteworthy:

      o An 18-year-old girl was bitten on her chin by a ferret being
      offered for sale in a shopping mall by an unknown free-lance
      vendor; he left with the ferret immediately after the incident.
      Rabies prophylaxis was discussed with the patient, but she was
      lost to follow up.

      o Reaching through the window of an auto parked in a shopping mall
      lot, a 23-year-old woman was bitten as she tried to pet a ferret
      held by its owner. The owner immediately drove away with the

      ferret.

      -9-
      o A 12-year-old boy was carrying his pet ferret in his partially
      opened jacket; the boy uttered a loud whistle, whereupon the
      ferret bit the boy's chest and neck.

      o Three adults were exposed to a recaptured pet ferret that
      subsequently developed rabies. Evidently, the infection was
      acquired from a skunk.

      No report of the circumstances attendant to the attack were given
      for 14 cases.

      Eight (57 percent) of the California infants and children aged 3
      or younger were bitten on the head (5), neck (2), or face and
      hands (1). The remainder (six) were bitten only on the limbs. All
      attacks involving the head or neck and three involving the limbs only
      were unprovoked. Sites of head wounds were specifically reported (with
      some overlap) as follows: head, two; face, three; cheek, one; and
      lips, one. The wounds of eight infants were described as multiple
      punctures or lacerations; two were described as single bites or
      lacerations; and four were reported as just "bitten." One 10-month-old
      infant had 20 lacerations and punctures on the right side of her neck.
      A two-year-old had four bites on the right side of her neck. A
      5-month-old girl had 50 puncture wounds on her face. All the
      victim-facilitated or provoked attacks consisted of only etngle bites
      or scratches.

      Medical treatment reports were received on only 5 of the 14
      infants. For these cases, plastic and reconstructive surgery was

      required for one. Rabies prophylaxis was also administered to one.

      -10-
      Among the older children and adults, three (five percent) were
      bitten on the head or neck. These were all victim-facilitated or
      provoked. As many as ten of the limb bites were unprovoked, as
      described above. Bites on limbs were usually singular.

      For bites on the extremities, the wound sites were: finger, 22 (2
      multiple); hand, 19 (4 multiple); wrist, 4; elbow, 1; arm, 1; toe, 3;
      foot, 2 (1 ferret bit both toe and foot); ankle, 2; Achilles' tendon
      area, 1; unreported, 13. Wounds were described as "bitten" in 37
      cases; "bite" in 12 (5 multiple); "puncture" in 12 (6 multiple);
      "laceration" in 4 (1 multiple); "scratch" in 2; preexistent scratches
      contaminated by saliva of a sick ferret in 1; and unknown in 2.
      Overall, about 22 percent of reports indicated multiple wounds.

      Relevant wound data were provided for only 3 of the 13 victims of
      unreported age. Two received finger bites (one multiple), and one
      received multiple hand bites.

      Medical treatment data were provided for only 27 of the 73 (37
      percent) victims over 3 years of age. Six (22 percent) of these
      persons received rabies prophylaxis.

      As above, reported data on ferrets that attacked persons over
      three years of age are given here by numbers of persons attacked to
      facilitate evaluation from the attack perspective and to avoid
      confusion due to multiple bite victims per ferret or vice-versa.
      Although most ferrets bit only one victim, six ferrets bit two victims
      each, and three ferrets bit three victims each.

      Twenty-seven (87 percent) of the attacks where information was

      reported, were by ferrets one year of age or older. Twenty-five (68

      —li=
      percent) were by male ferrets. Three (21 percent) attacks, out of 14
      having data concerning reproductive intactness, were from neutered
      ferrets. Of the 42 reports containing relevant information, all the
      attacks were inflicted by ferrets that lacked state licenses.

      Fifty-one (55 percent) of the 92 animals for which information was
      available were identified as pets; the remaining 41 (45 percent) were
      presumed to be stray or feral animals. Of the 51 attacks by ferrets
      reported to be pets, 48 (94 percent) were from ferrets with identified
      owners--14 (29 percent) of these were from pets owned by the victim's
      family, and 34 (71 percent) had other owners.

      Three (four percent) of the bites by ferrets that were tested for
      rabies were inflicted by a rabid ferret.

      Attacks on Persons in California, Oregon, and Arizona

      The Department's request for data in early 1986 resulted in
      maximal California reports for that and the two adjacent years, whereas
      data received from Oregon and Arizona were nearly all retrospective,
      being largely pre-1986. Moreover, Oregon data were unavailable for
      1980 and 1984, and that state's ferret bite reporting system had been
      rendered largely nonfunctional in 1984. Thus, data from the three
      states are both incomplete and somewhat staggered temporally (Table 4).
      Altogether, 100 reports were available from California, 76 from Oregon,
      and 249 from Arizona, for a total of 425 attacks for the 3 states,
      spanning the 10-year period 1978-1987. Most of the reported incidents

      occurred between 1980 and 1985.

      -12-
      Data collection was not uniform for the three states. However,
      data on ages of victims were compared as closely as report categories
      and judgment permitted (Table 5). Persons 4 years old or younger
      comprised 11 to 16 percent of victims, averaging 13 percent.

      Gross estimates of annual rates of ferret attacks on people were
      made for the three states (Table 6). Near maximal reported numbers of
      attacks per annum were assumed to be typical (since reporting is almost
      certainly more likely to be deficient than inflated), and were divided
      by the State's approximate population to obtain a rough estimate of the
      number of ferret attacks per million human residents. Of note, while
      the Arizona reports came only from Maricopa and Pima counties, these
      two counties contain most of Arizona's population.

      The estimated rates suggest that ferret attacks, per one million
      human residents, were 1 in California, 7.4 in Oregon, and 25 in Arizona
      (Table 6). Data from Oregon indicated that 36 percent of their attacks
      were from stray or feral ferrets; 45 percent of California attacks were

      from stray or feral ferrets. a

      Twelve cases of ferret rabies have been documented in the
      United States, six of them since 1985 (Table 7). It is either known or
      reasonably assumed that essentially all of these ferrets had been
      bitten by rabid wild animals. (It is possible that one ferret may have
      developed a live rabies virus vaccine infection. Another ferret
      apparently was infected by a 13th rabid ferret to which it had been
      bred.) One ferret fought a raccoon in an area noted for raccoon rabies

      shortly before it developed the disease. In two incidents, one of

      -13-

      L—
      which was in California, escaped ferrets developed rabies shortly after
      they were recaptured.

      Rabies-infected ferrets also have been purchased in pet shops. In
      one case reported to us, a child was bitten in the face by a rabid
      ferret purchased at a pet shop. In another case, a ferret died of
      rabies a month after its purchase from a pet shop.

      Feral Ferrets in California

      According to information provided by DFG, there is scant evidence
      of feral ferret populations in California, at present. However,
      several years ago, a ferret kitten was found near its mother after the
      adult had been hit by an automobile in Kern County. It was concluded
      that the female had bred in the area. Since then, a male and female
      pair of ferrets have been live-trapped at Folsom Lake (Placer County),
      and another pair was trapped in Sonoma County. Sightings of individual
      ferrets, made by knowledgeable and reliable observers, also have been
      reported from Sonoma, Napa, Riverside, and San Francisco counties. In
      the latter instance, the ferret was emerging from a burrow at
      Candlestick Point Recreation Area.

      Animal control personnel and county public health laboratory
      directors in Northern California frequently report observations and
      captures of single ferrets. It is usually impossible to distinguish
      escaped or released pets from feral animals, although the former seem
      more likely to be observed in populated areas. Similarly, ferrets that
      approach people, sometimes inflicting bites as persons pet them, are

      likely to be stray pets.

      -14-
      “ewer

      A.

      At this time, the available information suggests that if feral
      ferret populations exist in California, they may not yet be beyond
      control. It must be emphasized, though, that feral ferrets abound in
      other states with climates far more severe than occurs in most of
      California, and that California's poultry producing and game bird

      producing areas provide habitats especially attractive to ferrets.

      IV. DISCUSSION

      History of Ferret Selective Breeding and Interaction With Man

      Ferrets were developed by man from polecats, which have a
      reputation for being extremely bloodthirsty, killing far more than they
      can devour and indiscriminately attacking any and all animals within
      range (Johnston, 1903). “In addition to killing native animals,
      polecats also have been reported to ravage small livestock, such as
      rabbits and poultry. Larger animals also may be attacked. For
      example, it has been reported that 16 turkeys were killed during a
      single night by one polecat and 10 ducks by another, with each victim
      being left with a hole in its neck (Bell, 1837). Feral ferrets behave
      as polecats (Corbet and Ovenden, 1980), and have been known to engage
      in wholesale slaughtering of livestock (Everitt, 1897; Dolensek and
      Burn, 1976; Harding, 1915).

      The savage characteristics of polecats were highly valued and
      emphasized in man's selective breeding and development of ferrets for
      killing rats and rabbits or for driving them from their holes so they

      could be killed by men or dogs for sport or pest control (Everitt,

      -15-
      1897). Unusually large and persistent fighters also have been bred to
      attack skunks and minks (Harding, 1915).

      In addition to being developed to be bold, man selectively bred
      ferrets to be unafraid of humans in order to get the animals to return
      on command and to facilitate handling them (Thomas, 1946). Thus,
      unlike polecats, ferrets lack an innate fear of man (Poole, 1972). An
      interesting anecdote in this regard involved a case of four wild
      ferrets from a reproducing feral population in Washington State that
      were found feeding on the carcass of a cow when a veterinarian arrived
      to autopsy the animal. The ferrets refused to leave and continued to
      chew on the carcass throughout the autopsy (Porter, 1987).

      Although ferrets have been selectively bred over the centuries,
      one should not presume that man's development of ferrets from polecats
      means that the ferret has undergone isolation from and differentiation
      from polecats for thousands of years. On the contrary, ferret breeders
      have periodically crossed ferrets with polecats to produce the
      polecat-ferret or fitch-ferret coat color pattern (Fennell, 1841;
      Matthews, 1968; Corbet and Southern, 1977).

      Training of ferrets, as well as those who handle them, is required
      in order to prevent the animals from biting or attacking their
      handlers. Ferrets that have undergone training not to bite persons
      find a near parallel with other wild species that have received similar
      rearing or training. All such species are characterized by a
      relatively high frequency of biting compared to dogs, and some (e.g.,
      wolves, wolf-dog hybrids, coyotes, raccoons, and ocelots) have killed

      human infants and/or eaten their tissues (Constantine, 1986). However,

      -16-
      ferrets seem to present a somewhat different hazard, because, like pit
      bulls, they have been bred not only to be instinctively unafraid of man
      but also to be especially ferocious and tenacious against intended
      victims. Whereas trained ferrets may exercise restraint in biting
      adults, human infants may be regarded as prey to be killed and eaten.
      That ferrets are capable of restraining this behavior may be inferred
      from the data showing that these attacks have almost always occurred
      when adults were absent from small children. It is as if the animal
      waited until the adults left before attacking the hapless infant. Add
      to this the adeptness with which ferrets escape cages, and all the
      circumstances are present for tragedy to occur, as has been reported.
      Thus, it appears as if training of ferrets may not be entirely
      effective in preventing attacks on infants.

      For several reasons, infants may be somewhat irresistible to
      ferrets due to odors, sounds, taste, and actions resembling that of
      their natural prey. Ferrets encountering a nest of suckling rabbits
      underground will typically kill and eat them all, regardless of
      training (Brodie, 1978). Sucking or squeaking sounds that resemble
      utterances of rabbits in pain stimulate attacks by many kinds of
      predators, including ferrets. We believe that sounds made by human
      infants are among stimuli that trigger predatory ferret behavior.

      Bell (1837) cautioned persons against confidence in "the tameness
      of this sanguinary and deceitful animal" that had on multiple
      occasions attacked infants through "the resuscitation of its inherent

      though dormant propensity for blood." He declared:

      -17-
      "The Ferret exhibits a considerable degree of tameness, but

      without any discriminating attachment: it is in fact nothing

      more than the indifference and absence of fear and anger,

      which are the result of hereditary dependence upon and

      association with mankind. It will allow itself to be handled

      and played with, and in some cases may be suffered to run

      about the house with impunity, if it is carefully watched and

      well fed. But all this appearance of innocence and good

      temper is deceptive; and the Ferret, when tempted by

      opportunity, and excited by the smell or taste of blood,
      becomes as savage, and as indiscriminate in its attacks, as

      the Polecat itself.”

      Man's development of ferrets to be a more ferocious and effective
      predator than its polecat progenitor cannot be construed to be a form
      of domestication like that of animals designed to be pets, beasts of
      burden, or food animals. Indeed, even ferret proponents have not
      regarded ferrets as domestic animals; they have categorized them as
      less than domestic. Everitt (1897) quoted legal opinion that ferrets
      are not domestic animals, in contrast to animals generally acknowledged
      to be domestic. At a loss for adequate terminology, Harding (1915)
      stated that ferrets "are capable of only partial domestication,"

      cautioning that "they never cease to be dangerous if not carefully

      watched, especially where infants are within their reach."

      Problems Consequent to Pet Ferret Ownership

      Ferrets are purchased as pets because they are cute and playful.
      Owners usually give them the run of the house in order to play with
      them and enjoy their antics, although many owners confine them to
      indoor cages at night because they normally are most active at night.
      However, ferrets are very adept at escaping cages and households (e.g.,
      often through clothes dryer vents), and they can be difficult to find

      and recapture. Upon realization that one's pet ferret is missing, a

      ~18-
      major promoter of pet ferrets advises organizing a search of
      surrounding houses (Morton and Morton, 1985). Thus, if one owns a
      ferret, one can expect difficulties confining the animal and
      interaction of the animal with neighbors.

      Ferrets become especially bite prone when they reach about three
      months of age. Ferret play includes mock attacks, which in young
      animals are merely practice for adult behavior (Biben, 1982). People,
      particularly infants and strangers, may react to ferrets in a manner
      that encourages or exacerbates attack behavior.

      Attacks other than mock attacks can have a variety of causes
      (Fennell, 1841; Holz, 1982; Moody et al., 1985; Morton and Morton,
      1985; Roberts, 1977; Ryland et al., 1983; Wellstead, 1982; Willis and
      Barrow, 1971; Winsted, 1981). For example, circumstances observed to

      precede attacks include the following:

      Tpie>: Normal behavior. (As soon as young animals open their eyes, they

      TRUE

      «

      will treat as food anything put near their mouths until they learn
      better.)

      2. Lack of training not to bite. (Such training must be initiated
      very early, although some animals seem refractory to training, and
      training is not entirely successful in any case.)

      3. A new environment.

      4. A "nervous disposition" in the ferret.

      5. Anxiety or fear, i.e., anything that upsets the animal.

      «iS.
      6. Hunger. (Ferrets require a constant supply of food having a high
      content of animal protein because food transit time is only three

      hours.)

      FAISET. The smell or taste of blood.

      ? 8. Attempts to confine or Manipulate the ferret against its will.

      ? 9, Quick movements or sounds that startle the animal.

      tw ? 10. Pain.

      ll. "Natural aggressiveness" in uncastrated males.
      12. "Natural aggressiveness" in females from one week before

      parturition through lactation.

      FA/sE. 13. Mistaking as food fingers smeared with meat juices.

      FALSE 14. Anatomic parts, movements, or sounds that simulate the animal's

      natural prey.

      FFAlsE 15. Illness or disease (e.g., rabies).

      V he pe iS NO

      It is unclear why ferrets bite and sometimes hang on tenaciously
      to faces or limbs of persons who are sleeping or merely looking at,
      holding, or petting them. Likewise, it is unclear why these animals
      sometimes run to and reportedly viciously attack people on their lower
      limbs. Tenacious holding of the neck, with simultaneous blood

      drinking, has been reported in young ferrets upon the very first

      VAT URIL pred: {encounter with natural prey, suggesting that this is instinctive
      7k %,

      lomesr/e Fepeers

      be CAPTINE
      Ce lel sud ent

      MMCkCIAL Pet
      NOT E.KIST

      W The *witd.”

      behavior (Everitt, 1897). Perhaps, in some instances, unrecognized
      human activity has a triggering effect on such instinctive behavior.
      Stimuli that apparently incite ferrets to attack sleeping infants may

      include some of the previously mentioned circumstances or may be due to

      (2/T here. po The. identification of infants as prey.

      ERROTS

      -20-
      Bites to hands of persons who handle ferrets can be decreased by
      heeding commonly espoused advice to wear gloves when handling these
      animals (Rowlands, 1967; Willis and Barrow, 1971; Roberts, 1977;
      Winsted, 1981; Wellstead, 1982; Ryland et al., 1983), although holding
      a ferret with leather gloves heavy enough to fully protect against the
      animal's canine teeth may injure the animal (Hammond and Chesterman,
      1972).

      There are additional notable problems peculiar to ferret
      ownership. Ferrets have anal sacs and other cutaneous glands in the
      perianal skin that emit an obnoxious smelling substance, the production
      of which can be decreased but not entirely eliminated by "descenting"
      surgery similar to that performed on skunks (Creed and Kainer, 1981).
      Frequent bathing, especially of males, that removes greasiness from the
      coat also decreases the odor. Surgical neutering is said by some
      persons to lessen the tendency to bite, but others claim it does not
      help. Female ferrets are seasonally polyestrus, exhibiting signs of
      estrus from spring to fall, and they are induced ovulators. Unless
      they are bred or neutered, half of females will die from hypoplastic
      anemia (Ryland et al, 1983). These things, combined with the
      relatively short life span of ferrets (usually about half that of dogs
      or cats) are further reasons why ferrets would appear to be less
      desirable pets than other common pet species.

      Given the foregoing kinds of difficulties peculiar to ferret
      ownership, especially biting, many owners try to rid themselves of the

      animals by (1) returning them to pet shops or other source of

      -21-
      9o30 pith APBIES
      “7 - BlCAKS tr
      4. ~ LSUPLEY
      BITS, SKUMKS
      IND SGA(REELS

      ia
      )

      MOT ;

      YVERIFIFD. L

      acquisition, (2) selling or giving them away, (3) turning them loose,
      (4) turning them in to animal control authorities for destruction,
      often accompanied with requests for rabies tests, and (5) not
      endeavoring to recover escaped animals. Of course, once loosed into
      nature, ferrets are likely to interact with wild animals, and they may
      become infected with rabies. Whether or not they get rabies, under
      these circumstances they may bite other persons, including those they
      befriend, necessitating antirabies prophylaxis for the bite victim. In
      addition, released pets may initiate feral populations.

      Despite the difficulties of pet ferret ownership, many owners are
      exceedingly attached to their animals. For example, several ferret
      owners with extensively assaulted infant children apparently have
      seemed to place the welfare of their ferrets above that of their
      children, taking actions to prevent destruction of their ferrets that
      is required to perform rabies tests. In one recent case, a mother
      whose ferret had savagely attacked her newborn infant and attacked the
      face of a neighbor's infant succeeded in preventing destruction of the
      ferret, reportedly declaring she would rather destroy the child,

      because she had possessed the ferret for a longer time.

      Rabies in Pet Ferrets

      Twelve cases of rabies in pet ferrets are known to have occurred
      in the United States since 1958 (Table 7), ten of them in the last
      seven years. During this period great effort has been made by ferret
      proponents to market ferrets as pets. Available reports indicate that

      22 percent of Californians who were bitten by ferrets underwent rabies

      -22-
      prophylaxis. Other state's reports that were received indicate that
      the same proportion (22 percent) of assaulted infants and young
      children were given rabies prophylaxis. In a number of cases, though,
      the biting ferret was not available to be tested, making it necessary
      to administer rabies prophylaxis.

      No rabies vaccine trials have been performed in ferrets to justify
      licensing any product for that purpose; thus, no rabies vaccine can be
      guaranteed to be effective in ferrets. Moreover, some live virus
      rabies vaccines are known to produce rabies infections at unknown
      frequencies in wild carnivores, so live rabies virus vaccines should
      not be used in ferrets. False security could arise from use of killed
      vaccines not licensed for use in ferrets, generating health hazards and
      possible legal problems. -It is conceivable that trials with killed
      virus vaccines will be done and a product eventually approved for use
      in these animals. However, rabies vaccines used for dogs or cats,
      although tested and licensed, sometimes fail, especially when the
      vaccinated animal is permitted to run free and contact skunks or other
      wild animals that may transmit overwhelming doses of virus. Moreover,
      vaccines usually are prepared from laboratory strains of the virus,
      and these stock are known to differ somewhat from the many other
      strains found in wild animals, diminishing the vaccine's effectiveness
      in some instances (World Health Organization Expert Committee on
      Rabies, 1984). Given the adeptness with which ferrets escape and/or
      are released by their owners, thereby allowing contact with wildlife,
      vaccinated ferrets and their bites would still have to be regarded as

      potentially hazardous from a rabies standpoint. This concern is

      -23-
      especially significant in light of the number of reported ferret bites
      inflicted by stray or feral animals.

      A ferret that bites should be killed and tested for rabies as
      quickly as possible to determine whether the bite victim has been
      exposed to rabies (Centers for Disease Control, 1986b). However,
      because there is no optional grace period, it is probably prudent to
      initiate rabies prophylaxis immediately after the bite and later
      discontinue it if the animal proves not to be rabid (Public Health
      Service Advisory Committee on Immunization Practices, 1984). This is
      especially so for facial bites. To merely quarantine the animal and
      wait for it to develop rabies signs is problematic, for the maximal
      interval between an infectious bite and the advent of rabies signs in
      ferrets is unknown. Of interest, though, this interval is known to be
      as long as eight days in the striped skunk, another mustelid (Parker,
      1975). Also of concern, signs of rabies are little known in ferrets,
      and as such they may not be recognized. These signs are known to vary
      markedly in different species according to the viral strain and the
      dose received (Constantine, 1967).

      The claim that pet ferrets cannot get rabies because they are kept
      indoors is unsubstantiated, and is undermined by the frequency that
      these animals escape the home and by the details of known ferret rabies
      cases. Rabid ferrets have been encountered at the breeding or
      wholesale source, the pet shop or retail source, in the home, and in
      the wilds. It appears, then, that rabid ferrets may occur any place at

      any time.

      -24-
      D.

      Ferret Attacks According to Age of Victim

      As tabulated in Tables 2 and 3, the circumstances attendant to
      ferret attacks and the types of wounds inflicted differ according to
      the age of the person attacked. Infants aged three years or younger
      usually were attacked without provocation by ferrets known to be pets,
      and they generally were attacked when asleep or lying down. The wounds
      in these cases typically consisted of multiple bites or lacerations,
      usually to the head and/or neck. In contrast, older children and
      adults usually provoked or otherwise prompted the attack. About half
      of the attacks on these persons involved Stray or feral animals, with
      the victim being awake and ambulatory at the time of attack. The
      wounds in these cases typically consisted of single bites inflicted on
      the extremities. The differences in circumstances attendant to bites
      on infants and older persons and the differences in the anatomic site
      and extensiveness of the wounds were significant (p<0.01). In
      addition, the data suggested that older persons were more likely to be
      attacked by male ferrets, while attacks on infants involved male and
      female animals equally (p<0.05).

      The foregoing differences seem explainable largely on the basis of
      opportunity. A sleeping or reclining infant is far more susceptible to
      repetitive attack than are older, ambulatory persons. Likewise,
      infants are exposed primarily to pet animals, rather than strays.
      Similarly, male ferrets are reported to be more aggressive than females
      (Lavers, 1973; Biben, 1982), and probably are more inclined to approach

      or attack ambulatory persons.

      ~25-
      Several cases occurring outside of California are worth describing
      because of their unusual circumstances that contrast with the above
      noted general patterns of attack according to the victim's age. For
      example, a sleeping six-year-old Oregon girl was bitten on the face
      beneath her eye while sleeping (Williams, 1986a). A 17-year-old
      New York girl was also bitten on the face under similar circumstances
      (Orr, 1986). Whereas adults usually have no problem promptly
      terminating attacks, it is not always possible to do so. For example,
      an animal shelter employee in Nevada opened the door to a previously
      playful ferret's cage in order to move the animal to another cage, and
      the animal quickly ran up his arm, over his shoulders, and down the
      other arm, inflicting 20-30 bites along the way (Coffey, 1985b;
      Boneck, 1988). This was similar to an attack on a New York infant, in
      which case bites were inflicted at about one-fourth-inch intervals all
      along the arm. Two other cases of note involved a two-year-old Oregon
      boy who was bitten on the end of his penis while sleeping (Williams,
      1986a), and a seven-year-old Arizona boy who was similarly bitten
      (Wright, 1986).

      Another noteworthy incident involved a ferret sitting alongside a
      road at night. The victims, who were driving by, stopped and opened
      the front door of their car, whereupon the ferret hopped into the car
      and onto the front seat between the driver and his wife, who was bitten
      when she tried to pet the animal. The ferret would not release its
      bite grip, and the husband was bitten when he tried to force its
      release. Subsequently, he delivered the ferret to an animal shelter

      (Williams, 1986a).

      ~26-
      Ferrets Apparently Perceive Human Infants as Prey

      Observations consistent with the notion that ferrets perceive
      human infants as prey include the small size and helplessness of the
      victims, the anatomical locations of and extensiveness of inflicted
      wounds, and ferret ingestion of the tissues of live, human infants.
      Victim odors, sounds, and reactions to the initial bites may play a
      role, as may the taste of blood and flesh. The smell or taste of blood
      is said to stimulate savage and indiscriminate attacks by ferrets
      (Fennell, 1841).

      Our data show that attacks were more frequently reported in
      infants less than a year of age, decreasing with increasing infant age
      and size. Apfelbach and Wester (1977) reported that ferrets are
      inhibited from attacking pexceived ‘prey as the latter approach more
      than double the size of the ferret. The same authors reported that
      when hunting, ferrets appear to innately aim at the most anterior part
      of the prey, which is consistent with the extensive head and neck
      attacks involving infants.

      That the attractiveness of infants as victims is not due solely to
      their small size is suggested by a 1987 New York incident wherein a
      blind, seven-year-old spina bifida victim received numerous leg bites
      and scratches during a nocturnal attack by two pet ferrets (Abelseth,
      1987). The inability of the victim to terminate an attack may

      encourage more extensive damage.

      -27-
      Suckling rabbits are among favored ferret foods in the wild, and
      certain characteristics (e.g., odors or sounds) shared by suckling
      humans may make the latter especially attractive "prey." As previously
      noted, certain sounds have been observed to trigger ferret attacks,
      with persons having unintentionally triggered seemingly instinctive
      attacks on themselves by ferrets. For example, while lying down
      outside a rabbit burrow, a ferret handler made squeaking and sucking
      sounds resembling utterances made by rabbits in pain. This
      successfully called his ferret out of the burrow, but the animal fixed
      its teeth firmly in the man's cheek after rushing from the burrow
      (Everitt, 1897). The same author referred to a ferret that nearly bit
      off its trainer's ear lobe. It also seems that ferret attacks may be
      triggered by certain sucking or squeaking sounds, including infants
      sucking on milk bottle nipples or pacifiers. However, the association
      of such sounds with prey may have to be learned in nature and, thus,
      may not apply to animals reared in captivity.

      Movements of various kinds are also known to elicit attacks. For
      example, the carcass of a dead rabbit may be ignored by a ferret but it
      has been observed to be attacked if it is shaken as if it were alive
      (Thomas, 1946). Movements of infants may also serve as triggers for
      attacks, especially quick or jerky movements.

      Ferrets and polecats seem to display "frenzy" behavior, such as
      occurs with sharks, during some attacks. In such situations they seem
      to become detached from any semblance of reason, functioning on a
      purely instinctive level. This may have survival value in nature.
      Illustrative of such behavior are reports of mass killings of chickens,

      *

      -28-
      ducks, turkeys, and rabbits in poultry houses or rabbit hutches. In
      these cases, ferrets have reportedly killed all animals present,
      although eating relatively little of the prey. In nature, ferrets
      store small game food (e.g., frogs or fish) in their dens, but they
      are generally unable to transport or store great numbers of larger prey
      such as chickens or rabbits.

      Also supportive of the notion of instinctive "ferret frenzy" is
      the seemingly unexplainable situation of a ferret seizing a familiar
      and friendly adult person with its teeth and then refusing to let go,
      especially when the attack is triggered by simulated utterances of a
      rabbit in pain. Of interest, polecats commonly bite prey with such a
      "death grip,” although young, inexperienced polecats may grip the wrong
      site--perhaps due to their relatively poor eyesight (Southern, 1964).
      Similar simulated sounds have been used by hunters to trigger charges
      by large wild canids and felids. It is possible, then, that with
      suitable stimulation, the urge for ferrets to attack infants may be
      guided by instinctive behavior, especially when the inhibitory presence
      of adults is removed.

      A brief review of methods of ferret attacks on natural prey is
      relevant here. Upon recognizing prey, polecats and ferrets utilize
      typical musteline attack methods. The attacking animal leaps on the
      prey animal, clutching the victim's body with the forelimbs, and
      killing it with either a bite on the occipital region, which usually
      crushes the back of the skull (Jennison, 1927; Apfelbach and Wester,

      1977), or by a bite of the neck, which opens major blood vessels of

      -29-
      larger or long-necked prey (Ewer, 1973). The "death grip" to the head
      or neck is concomitant with violent shaking of the prey by polecats
      (Southern, 1964). The neck bite, often used to kill rabbit sucklings,
      is said to be instinctive but only partially developed in young
      polecats, requiring perfection by practice (Corbet and Southern, 1977).
      This seems to be accomplished, in part, by mock neck biting during play
      and by sustained neck biting during intraspecies combat (Biben, 1982).

      A further possible explanation of the repetitive and mutilating
      Nature of ferret attacks on infants is found in the older literature on
      ferrets. That is, ferrets may inflict hundreds of bites on infants to
      increase the flow of blood on which to feed. Similarly, they may
      extensively bite and tear the scalp and face in an effort to get at and
      eat brain tissue, as reported in musteline attacks on natural prey.
      Frequent reference is made in older publications of the marked appetite
      of ferrets, polecats, and other mustelines for the blood and brains of
      their victims. Everitt (1897) stated that lactating ferrets require
      blood or they will eat their offspring. Fennell (1841) and Roberts
      (1977) reported that ferrets drink the blood of their victims, and
      Harding (1915) claimed that in an abundance of slaughtered prey,
      ferrets merely suck the blood of their victims, a practice also
      attributed to polecats (Bell, 1837; Johnston, 1903). Bell, like other
      early writers, cited the victim's brain as the first choice of solid
      tissues eaten after kills made by polecats and other mustelines. The
      accuracy of such accounts cannot be verified, but the repeated mention
      of such behavior suggests that these tendencies be considered when

      analyzing attacks of ferrets on human infants.

      -30-
      Of further interest in this regard is an account of a pet ferret
      drinking human infant blood in England more than 150 years ago. Jesse
      (1834) wrote:

      "Some few years ago, a poor woman, holding a mangled infant in
      her arms, rushed, screaming with agony and fright, into my

      — Wh friend's house, who is a surgeon, imploring him to save the
      AW yo child's life, who, she said, had been almost killed by a
      Qo ferret; the face, neck, and arms, were dreadfully lacerated,

      ae Ww the jugular vein had been opened, as also the temporal
      \ A artery; the eyes were greatly injured, and indeed the child,

      Ve Ww who is still living, has lost the entire sight of one of
      ae a them, and has very imperfect vision in the other. Having
      stopped the still bleeding vessels, my friend accompanied the

      N) ny u mother to her cottage, on entering which the child, in some
      Y degree recovering from its state of apparent death, began to
      Ye cry, when the Ferret was in an instant seen rushing from

      behind some bavins where he had taken shelter, and, with his
      5 head erect, boldly came forward and met the infuriated parent
      eae We in the middle of the room, still holding the infant in her

      ¢ RY arms. On my friend's kicking the Ferret, as the first
      \ vy impulse of protection, the animal endeavored to seize his
      an ¢ leg, and not until his back was broken by repeated kicks, did
      ye 0 he give over his earnest and reiterated attempts to renew his
      ee oy sanguinary feast; and indeed, whilst in the agonies of death,
      a wt the piteous screams of the child seemed to rouse him to vain
      « efforts to regain his prey. The Ferret was of large growth,

      and much distended with the infant's blood; and although
      formerly of peculiar shyness, yet he lost sight of fear, and
      became ferocious in the pursuit of the unfortunate infant.
      It appears the poor woman had left her child (about six
      months old) in a cradle whilst she went to market, when it is
      supposed the infant's cry had arrested the attention of the
      Ferret, who managed to make his escape, and thus effected his
      purpose. There is good reason to believe he must have past
      more than half an hour in the indulgence of his appetite,
      from the circumstance of the neighbors having heard the
      piercing shrieks of the child for a long time without the
      slightest suspicion of the mother's absence."

      F. Liability of Ferret Ownership

      We are aware of litigation against pet shops and ferret owners
      consequent to ferret attacks. Therefore, in our review of these cases

      we tried to assess the nature of ownership of ferrets that attacked

      -31-
      persons. Based on the available information, it appears that 55
      percent of the ferret attacks in California were inflicted by known
      pets, with the remainder presumably due to strays or feral animals-~-at
      least they were of uncertain ownership. Of note, the ferrets of
      unknown origin may well have been former pets or strays because they
      did not hesitate to approach people. In fact, some were suspected to
      be pets of neighbors who were known to own ferrets, although ownership
      was typically denied following attacks. (If one were to consider the
      attacks of animals having unknown or disputed origin to be from pets
      of unknown owners and include them with the aforementioned attacks from
      pet ferrets not owned by the victim's family, then 84 percent of ferret
      attacks on Californians were inflicted by pet ferrets belonging to
      persons other than the victim, a percentage similar to what has been
      observed for dog bites.) Of the attacks from known pets, 29 percent
      were from ferrets residing in the victim's household.

      Data relevant to the 64 unprovoked attacks on infants are similar
      to the foregoing except fewer of the infants were outside and available
      to be attacked by stray ferrets. Overall, in 45 (70 percent) of these
      cases information on animal ownership was known. Of the attacks on
      these infants from known long-established pets, 39 percent were from
      ferrets owned by the victim's family and 61 percent were from ferrets
      of other known ownership. Of the 7 attacks that were inflicted on
      infants by stray ferrets, 6 (86 percent) were from animals that had
      just been brought into the household because they were friendly. Of

      the 21 known owners of attacking ferrets from without the victim's

      -32-
      family (i.e., for whom specific data were available), 7 (33 percent)
      were visitors, 6 (29 percent) were babysitters, 5 (24 percent) were
      hosts, and 3 (14 percent) were vendors.

      Our data indicate that 28 percent of attacked infants 3 years of
      age or younger required plastic and reconstructive surgery, especially
      of their faces. Some of the infants were left with permanent
      disfigurement. Four percent of bitten persons were exposed to rabid
      ferrets, and 22 percent of attacked persons were given rabies
      prophylaxis. It is possible that the actual occurrence of these
      sequelae are exaggerated, since more serious cases tend to be reported.
      However, it is also known that rabies prophylaxis is significantly
      underreported.

      Given the above findings and concerns, it appears that ownership

      or selling of pet ferrets may present significant liability risks.

      Feral Ferret Populations Can Devastate Small Livestock and Native

      Animals and Contribute to the Wildlife Rabies Problem

      Intentional or accidental releases of pet ferrets have led to the
      development of feral ferret populations. This continues to be a
      problem. Indeed, "throughout the centuries the number of ferrets that
      have escaped and returned to the wild must be enormous" (Matthews,
      1968). Such populations, a few of which have been trapped to
      extinction, have been reported in numerous places, including England,
      Wales, and Scotland (Corbet and Southern, 1977; Corbet, 1980; Howes,
      1980), mainland Europe (Lyneborg, 1971), Eurasia (De Vos et al., 1956),

      the Mediterranean islands of Sardinia and Sicily (Corbet, 1980; Konig,

      -33-
      1973), New Zealand (Corbet and Southern, 1977; Nowak and Paradiso,
      1983), and the United States (Dolensek and Burn, 1976; Stevens, 1979;
      Hoffmeister, 1986; Olson, 1987). As already noted, feral populations
      seem to also have developed in California in recent years, even though
      the animals are officially banned from the State.

      Feral ferrets are well known to behave like their polecat
      progenitors (Corbet, 1980), so it is not surprising that they have
      proved to be exceedingly destructive to small livestock like poultry
      and rabbits, as well as to native vertebrates. Their sometimes wanton
      destruction of small animals far beyond their food needs wreaks havoc
      on victimized native populations. Not only may adult animals be
      killed, but native species populations may be further harmed by the
      ferret's particular appetite for and ability to reach nestling mammals,
      birds, and eggs of the latter (ground-nesting species in particular).

      Long known for their potential to harm small domestic animals and
      wildlife in Britain and Europe, ferret populations were established in
      New Zealand to lower populations of rabbits that had been introduced in
      1864. Ferrets, and other introduced predators, now feed on native
      animals and have contributed to the extinction of 20 species of endemic
      New Zealand birds and have pushed many others to the brink of
      extinction (King, 1984).

      Dolensek and Burn (1976) report that European ferrets were
      imported into the United States about 1875 to kill rats. New London,
      Ohio used to be known as "Ferretville, U.S.A." due to its large-scale

      breeding and sales of ferrets. However, feral populations developed,

      ~34-
      and the animals became a plague to poultry producers. Subsequently,
      control measures became necessary, including passage of anti-ferret
      regulations by many states.

      Presently, a feral ferret population exists on San Juan Island,
      Washington, where the animals feed on introduced European rabbits,
      native animals, and possibly barnyard fowl. In addition to devastating
      European rabbit populations, they are believed responsible for
      reducing the native mink population in this area (Stevens, 1979).
      Similarly, at least three widely separated feral populations of
      European ferrets have been reported to be subsisting on prairie dog
      colonies in New Mexico (where they are competing with the near-extinct,
      native black-footed ferret) and have been reported to have attacked
      domestic poultry elsewhere (Olson, 1987). In a similar vein,
      Hoffmeister (1986), reporting on European ferrets living in the wild in
      various places in Cochise County, Arizona, has raised ecological

      concerns about feral ferrets in this area.

      Ferrets and Rabies

      As already discussed, ferrets can be infected with rabies
      transmitted to them by wild animals, and they can expose people and
      other vertebrates to the virus. Their ability to spread the infection
      among themselves is demonstrated by the infected caged ferret whose
      only known source of exposure was a ferret to which it had been bred.
      Moreover, rabies is regularly reported in European polecats, where
      populations of these animals have long been actively suppressed because

      of their destructive effects on livestock. Especially dangerous rabies

      -35-
      problems may develop in feral ferret populations in areas not native
      to the polecat but having rabies endemic in other species.

      There are two notable examples of outstanding rabies problems that
      have developed and persisted or grown worse in populations of
      introduced, nonnative carnivore species. At the turn of the century
      the raccoon dog was introduced from Korea into the far eastern Soviet
      Union, from which it has spread westward to Finland, Sweden, and
      Germany, becoming second only to foxes as a rabies vector in this part
      of the world (Kaplan, 1985). Similarly, Indian mongooses were
      imported to various Caribbean Islands in the late 1800s to control
      introduced pest rats and native poisonous snakes, but after destroying
      the rats they soon became the worst of all pests, greatly reducing the
      native fauna (i.e., all but exterminating several native species of
      mammals, birds, and reptiles) and causing serious economic
      repercussions by killing small domestic animals. Furthermore, the
      mongoose populations became infected with rabies, evidently from
      domestic dogs, and mongooses are now the most prominent rabies vector
      in the Caribbean Islands (De Vos et al, 1956; WHO Expert Committee on
      Rabies, 1984).

      Being fearless, savage and tenacious, ferrets may readily suppress
      native competitors and thrive on smaller native and domestic
      vertebrates, such as those found in California. The same
      characteristics should make them exceptionally effective transmitters
      of rabies among themselves and to other wild and domestic mammals,
      and to man. The presence of rabies in wildlife in most of California's

      counties constitutes a ready source of infection. The potential for

      -36-

      such transmission was realized in 1985, when an escaped pet ferret
      developed rabies of skunk origin upon recapture near Red Bluff,

      Tehama County, California (Table 7).

      VY. CONCLUSTON

      The legalization of ferrets as pets in California is opposed for
      Many reasons, as discussed in the preceding pages. Indeed, after
      reviewing the history of ferrets and the data gathered here, there
      seems to be no conclusion other than ferrets are miscast as pets. Even
      though some pet ferret owners are willing to suffer bites as a price of
      pet ownership, it is not reasonable to expect their neighbors and other
      persons to do so, especially in light of the potential for devastating
      attacks on infants and concerns about rabies, to say nothing of
      potential establishment of feral populations and the destructive
      effects on wildlife and small livestock. One might also question the
      humaneness of producing and marketing as pets any animal that typically
      must undergo two surgical operations in an effort to decrease biting,
      minimize offensive odors, and prevent death from anemia.

      Although ferret producers, vendors, and owners have been
      systematically pressuring various states and local jurisdictions to
      drop legal restrictions against pet ferrets (e.g., Anonymous, 1986),
      the converse has been encouraged by various organizations with
      knowledge of the hazards attendant to legalization of these animals as

      pets. Among the organizations opposing ferrets as pets are The Humane

      -37-
      Society of the United States (Herbet, 1987), The Defenders of Wildlife
      (Spotts, 1985), The Council on Public Health and Regulatory Veterinary
      Medicine of the American Veterinary Medical Association (American
      Veterinary Medical Association, 1985), the Council of State and
      Territorial Epidemiologists (Freeman, 1988), the United States Animal
      Health Association (Diesch, 1982b), and the Centers for Disease
      Control, United States Public Health Service (Centers for Disease
      Control, 1986b). To help translate the concerns into action, the
      Humane Society of the United States (1985) has developed model state
      legislation that restricts ferrets as pets.

      The differences in reported ferret attack rates between
      California and adjacent states (Table 6), where pet ferrets are
      unrestricted, indicate that California should continue to prohibit pet
      ferrets, albeit by no means entirely successful in this regard.

      If California's present estimated annual ferret attack rate of 1
      per million humans were to rise to that of Arizona, California's
      ferret-associated health problem would increase 25 fold. The increase
      might translate into annual increases in ferret attack reports on
      persons of all ages from 25 to 625 per year, rabies prophylactic
      treatments from 5.5 to 138 per year, and known exposures to rabid
      ferrets from 1 to 25. In addition, there would undoubtedly be a
      corresponding increase in financial costs for associated medical
      treatment and litigation. Likewise, the adverse effects of feral
      ferret populations on small livestock and wildlife would be increased.

      Enforcement of California's ferret exclusion rule is necessary.

      And while the ferret industry claims that hundreds of thousands of

      -38-
      ferrets are already in California illegally, that claim has not been
      verified, although there is evidence that efforts to import ferrets
      into California have increased greatly in the past few years. Ferret
      interceptions at agricultural inspection stations increased steadily
      from none in fiscal year 1975-76 to 210 in 1985-86 (California

      Department of Food and Agriculture, 1987). Greater surveillance and

      enforcement efforts in this regard may be needed.

      -39-
      Neva

      CASE

      NO. YEAR SITE
      1. 1978
      2. 1978
      3. 1978
      4. 1978
      5. 1978
      6. 1978
      7. 1978
      8. 1980
      9. 1981

      Table 1

      Unprovoked Ferret Attacks on Infants and Young Children

      VICTIM*

      INJURY

      London, 6mo F
      Eng] and

      Maryland Sday M

      South Child M
      Carolina

      Oregon 6yr F

      16mo

      Ohio

      ‘Arizona Child

      Exsanguin-
      ation due
      multiple
      bites of
      face

      Multiple
      bites of
      head, face,
      neck and
      arms

      Face bite,
      Antirabies
      treatment

      Cheek bite

      No bite;
      attack
      interrupted

      Scalp bite

      Cheeck bite,
      had to pry
      loose

      "Mauled, u
      including

      CIRCUMSTANCES
      OF ATTACK

      2 ferrets escaped
      cage, entered baby
      carriage in bedroom
      where infant was
      lying

      Ferret escaped cage,
      entered crib and

      attacked sleeping
      infant

      No details
      available —

      Child in bed, father
      bitten trying to
      remove ferret

      Above ferret ran,
      jumped into crib;
      father bitten

      Infant asleep in
      playpen

      Ferret dead next
      day

      Attack initially
      unobserved, then

      face; massive interrupted

      hemorrhage

      Multiple

      puncture
      wounds of

      finger

      Crawled on child

      SOURCE
      OF
      FERRET

      Family
      pets

      Visitor’s
      pet

      Purchased
      same day

      Family
      pet

      RABIES
      TesT**

      Pos

      REFERENCES

      Anon 1978, 1979
      Diesch 1981, 1982a

      Lebar 1978;
      Price & White 1978;
      Friedland 1978

      Diesch 1981

      Williams 1986a

      Williams 1986a

      Williams 1986a

      Williams 1986a

      Stevens 1980

      Wright 1986
      10.

      ll.

      12.

      13.

      14.
      15.

      16.

      17.

      18.

      19.

      2l.
      22.

      198]

      198]

      1981

      1981

      1981
      198]

      1981

      1982

      1982

      1982

      1982

      1982
      1982

      Arizona

      Arizona

      Arizona

      Arizona

      Arizona

      Infant

      Child

      Smo M

      Colorado 5mo F

      Nevada

      Arizona

      Arizona

      Arizona

      Arizona

      Arizona

      Arizona

      7mo M

      Child

      Child

      Child

      3wk M

      amo F
      2yr M

      Multiple Crawled into crib

      puncture
      wounds and

      laceration
      of face

      Single lacer- Crawled on child
      ation (bite)
      on leg

      Multiple
      lacerations
      on cheek,
      wrist, leg

      Baby sleeping

      Multiple

      puncture
      wounds of

      arm and leg

      Baby sleeping

      Forearm bite At relative’s hame

      40% ears Infant attacked in
      eaten; crib

      multiple face

      bites :

      Hundreds of Infant asleep in
      bites on in crib; ferret was

      face, hand, 4mo M, not neutered
      back of knee

      Multiple
      puncture
      wounds and
      laceration
      on neck

      Child sleeping

      Multiple

      puncture
      wounds of
      scalp & lip

      Child sleeping

      Multiple
      lacerations
      above eye

      Child sleeping

      Wrist
      "bitten"

      Diapers being
      changed
      Arm “bitten" At relative’s hone

      Face Unknown

      "bitten"

      4]

      Host’s pet

      Found in
      yard day
      before

      Pet of
      babysitter

      Family
      pet

      Family
      pet

      Neg

      Neg

      Wright 1986

      Wright 1986

      Wright 1986

      Wright 1986

      Wright 1986
      COC 1981

      Coffey 1985a, 1985b

      Wright 1986

      Wright 1986

      Wright 1986

      Wright 1986

      Wright 1986
      Wright 1986
      23.

      24.

      25.

      26.

      27,

      30.

      31.

      32.

      33.

      1982

      1983

      1983

      1983

      1983

      1984

      . 198

      1984

      1984

      1985

      1985

      Maryland 2mo M

      Arizona amo F

      Nevada 50 M

      Oregon limo M

      Arizona Child

      Arizona Child

      Cali- l0mF
      fornia

      Arizona 5mo M

      Cali- &oM
      formmia

      Face Unknown
      "mauled";

      antirabies

      treatment

      Bites to foot Crawled into baby’
      and ankle bed

      Hundreds of Infant asleep on
      of bites to bed; 7mo unspayed
      to face, ferret escaped
      eyelids, cage

      back of head

      hands, wrists

      4)

      Bites on Attacked infant in
      head and crib
      face

      Bite on hand Infant lying on
      floor

      Multiple Child asleep
      puncture

      wounds on

      cheek

      Single Child asleep

      puncture
      wound on face

      20 lacera- _‘ Ferret escaped cage,

      tions and attacked infant in

      puncture crib; ferret was
      wounds on uncastrated 7-9n0 M
      right side

      of neck

      Severe, Ferret jumped

      multiple into face of
      bites on child while
      face; had to she was on
      pry animal _— the toilet

      off cheek

      Bites on Ferret entered crib

      back of

      neck, fore-

      head, hands

      Bite on Infant attacked

      hand while lying on a
      blanket on the
      floor

      42

      Non-family
      pet; owner
      would not
      sacrifice

      Pet of Neg
      babysitter

      Pet shop --

      Visitor’s --
      pet

      Illegal pet

      Pet having
      several
      previous
      owners

      Illegal pet Neg
      (obtained 2
      days earlier
      from owner

      of 2 yrs)

      AP 1982

      Wright 1986

      Coffey 1985a, 1985b

      Williams 1986a

      Williams 1986a

      Wright 1986

      Wright 1986

      Kelly 1986

      Williams 1986a

      Wright 1986

      Miller 1986
      35.

      36.

      37.

      39.

      41.

      42.

      43.

      1985

      1985

      1985

      1985

      1985

      1985

      1986

      1986

      1986

      1986

      1986

      Cali-
      fornia

      Indiana

      Virginia 20m F

      Nevada 29da F

      North
      Carolina

      North

      Carolina

      Arizona

      Cali-
      fomia

      Cali-
      fornia

      Cali-
      fornia

      Cali-
      fornia

      18m0 F Bite on head; Unknown

      &mo

      lmo M

      Smo

      Baby

      Smo F

      3yr M

      1&no F

      Antirabies
      treatment

      Extensive
      bite wounds
      of the face,
      hands and
      ayms

      Bites on
      ankle.

      Nose eaten;
      chewed eye-
      lids, lips,
      face and
      hands

      Ear eaten

      100 puncture
      wounds to
      face and head

      Multiple

      puncture
      wounds of

      wrist

      50 puncture
      wounds of
      face

      Bites &
      scratches
      on hand &
      feet

      Bites on
      face

      Multiple
      bites on
      face

      Ferret escaped cage
      and attacked infant
      in crib

      2yr F ferret
      escaped cage,
      ran to and bit
      child walking
      with mother
      near the
      ferret’s home

      Attacked infant at
      2 a.m. while child

      sleeping in playpen;
      ferret uncastrated
      5mo M

      Ferret attacked
      infant in bed

      Unknown

      Crawled on baby

      Infant on floor with

      milk bottle when
      attacked

      Ferret found near
      screaming child

      Not reported

      Not reported

      43

      Ferret owner --
      was visitor
      who left

      with animal

      Caught Neg
      in yard

      Belonged Neg
      to another

      party

      Pet Neg

      Pet --

      Family --
      pet

      legal Neg
      pet

      Feral or Neg
      stray

      I] legal
      pet (Feral
      or stray
      ferret
      adopted by
      family

      Illegal
      pet

      Fisher 1986

      AVA 1985;
      Diesch 1986

      Anon 1985;
      Carton 1985

      Coffey 1985a,1985b

      Freeman 1987

      Freeman 1987

      Wright 1986

      California
      Department of Food
      & Agriculture 1986;
      Tacal 1987a

      Weeks 1986

      Giles 1986

      Avedian 1986
      45.

      47.

      49.

      51.

      52.

      55.

      1986

      1986

      1986

      1986

      1986

      1986

      1986

      1986

      1986

      1986

      1987

      Cali- 14m M

      fornia

      Jersey

      Mexico

      New 3yr M
      Mexico

      Texas

      Texas Infant

      Wash- 180
      ington

      New York 9mo F

      Wash-
      ington

      2da F

      Bites on
      lip, cheek,
      and hands

      20-25 bites
      on arms and

      legs, deep

      laceration

      on foot sole

      Bitten on
      face

      Bitten on
      face

      Over 200
      bites on

      cheek, hands
      forearms and

      back

      Bit end
      of penis

      Severe bites Ferret escaped cage,
      and scratches entered infant’s
      crib during the

      to ears,
      around eyes
      and arm

      Severely
      bitten on
      face and
      ears

      Severely

      bitten on am
      & leg; would

      not let go

      Multiple
      bites of
      hand and
      forearm

      Ferret attacked
      child in bed

      Attacked in crib at
      babysitter’s home

      Infant in crib

      Child in crib

      Infant sleeping in

      crib

      Climbed under bed-
      covers with sleeping
      ‘child

      night

      Uncaged ferret
      attacked infant in

      crib

      Unknown

      Ferret attacked
      infant sitting on

      the floor

      Ferret entered
      bassinet; mother
      awoke observing
      ferret with its teeth
      in scalp flipping
      infant over "like

      a piece of meat."

      4

      Illegal
      pet

      Babysitter’s Neg

      pet for 2yrs

      Family
      pet

      Feral or
      stray,
      brought
      into hore
      Family pet
      for 6 mo

      Pet (pur-
      chased as
      kit 3 mo
      earlier)

      Family pet
      for 1 yr

      Family
      pet

      Family
      pet for
      4 m

      Family
      pet

      Host’ s
      pet

      Neg

      Tacal 1987b

      Sorhage 1986, 1987

      Hull 1986

      Hull 1986

      Williams 1986b

      Williams 1986a

      Clark 1987a, 1987b

      Clark 1987a

      Nicola 1986

      Abelseth 1987

      Gilmore 1987
      57.

      59.

      61.

      62.

      1987

      1987

      1987

      1987

      1987

      1987

      1987

      1987

      1987

      Texas

      Texas 14da

      Cali-

      fornia

      5mo F

      Cali-
      fornia

      5mo F

      Colorado 4mo F

      New 4mo
      Hampshire

      Wis-
      consin

      7mo F

      New York 6mo F

      Cali-
      fornia

      ayr F

      Facial bites Ran to and attacked Pet; _

      Bitten
      many times
      on arms

      Multiple
      bites on
      forearm
      and elbow

      "Wounds
      to head"

      Multiple
      puncture
      wounds and
      lacerations
      to face,
      including
      conjunctiva

      Bites to face 2 ferrets escaped
      and hands cage; entered crib

      9 bites to
      scalp &

      3 scratches
      from eye to
      tenple

      Unknown

      4 bites on
      right side
      neck; had

      child on the floor;
      unspayed F ferret

      Above ferret entered

      crib of sleeping
      infant

      Unspayed, 2yr F

      ferret climbed into
      bassinet to attack

      Ferret attacked

      child in bed; attack

      interrupted

      Ferret escaped cage;

      attacked child

      Child playing on
      floor when attacked

      Ferret entered crib

      were infant was
      lying

      Asleep in crib at
      babysitter’s home
      when uncaged ferret

      to pry loose attacked

      Purchased at
      pet shop 2
      yrs earlier
      by another
      conmune
      member

      Same as -
      above

      Family -
      pet for
      4 mo

      Babysitter’s --
      pet in
      owner’s hone

      Babysitter’s --
      pet in
      ower’s home

      Pets of
      visitor

      Pet Neg

      Visitor’s Neg
      pet

      Stray F Neg
      ferret

      adopted

      1.5 yrs

      earlier

      NOTE: The same amount of information is not available on all cases.

      * --M = Male; F = Female; yr = year; mo = month; wk = week; da = day

      wk -- Pos = Positive; Neg = Negative; -- = no information about

      child, baby and infant = young child of unspecified age
      but in all but one case appeared to be three years of age

      or younger.

      test results or no test performed.

      45

      Rutty 1987

      Rutty 1987

      Minor 1987

      Tacal 1987b

      Pape 1987

      Mock 1987;
      Clayton 1987

      Kurth & Weiss 1987

      Barr 1987

      Liska 1987
      TABLE 2. CHARACTERISTICS OF 64 UNPROVOKED FERRET
      ATTACKS ON INFANTS AND SMALL CHILDREN*

      Characteristics of Attack Number (Percent)

      Sex of Person Attacked

      Male 22 (48)
      Female 24 (52)
      Not reported 18
      Site of Wounds
      Head 30 (48)
      Neck 3 (5)
      Head or neck and limbs 11 (18)
      Limbs 18 (29)
      Assault blocked 1
      Site of wounds not reported 1

      Treatment Reported

      Reconstructive surgery 5 (28)
      Wound debridement, dressings, etc. 7 (39)
      Antirabies prophylaxis 4 (22)
      None 2 (11)
      Treatment not reported 45
      Characteristics of Attacking Ferrets

      Less than one year old 7 (39)
      Adult 11 (61)
      Not reported 47

      Male 7 (50)
      Female 7 (50)
      Not reported 51

      Pet 45 (87)
      Stray 7 (13)
      Not reported 12

      Neutered

      Not neutered 8 (100)
      Not reported 56

      Positive Rabies Exposure (FRA) 1 (4)
      Negative Rabies Exposure (FRA) 24 (96)
      Not reported 39

      * In all but one case, the children were three years old or younger.

      -46-
      TABLE 3. CHARACTERISTICS OF 100 FERRET ATTACKS ON CALIFORNIANS

      AGE OF PERSON ATTACKED

      _ Characteristics 3 Years or Younger >3 to 10 Years > 10 to 20 Years More than 20 Years Not Reported Totals
      aaa Number (Percent) | Number (Percent) | Number (Percent) | Number (Percent) | Number (Percent) | Number (Percent)
      Sex of Person Attacked
      Male 4 (29) 4 (50) 8 (62) 26 (59) 2 (50) 44 (53)
      Female 10 (71) 4 (50) 5 (38) 18 (41) 2 (50) 39 (47)
      Not reported 1 7 9 17
      Total persons attacked 14 9 13 51 13 100
      Site of Wound
      Head 5 (36) 2 (15) 7 (9)
      Neck 2 (14) 1 (8) 3 (4)
      Head or neck and limbs 1 (7) 1 (1)
      Limbs 6 (43) 9 (100) 10 (77) 38 (100) 3 (100) 66 (86)
      Not reported 13 10 23
      Treatment Reported
      Reconstructive surgery* 1 (20) 1 (3)
      Wound debridement,
      dressings, etc.* 2 (40) 2 (67) 2 (40) 7 (37) 13 (41)
      Tetanus prophylaxis 1 (20) 1 (20) 7 (37) 9 (28)
      Antirabies prophylaxis 1 (20) 1 {33) 4 (21) 6 (19)
      Tetanus and antirabies 1 (20) 1 (3)
      None 1 (20) 1 (5) 2 (6)
      Not reported 9 6 8 32 13 68
      Characteristics of
      Attacking Ferrets
      ' ess than one year old 1 (20) 2 (40) 1 (20) 4 (13)
      jult . 4 (80) 3 (60) 4 (80) 12 (100) 4 (100) 27 (87)
      “wot reported 9 4 8 39 9 69
      Male 3 (60) 2 (100) 3 (75) 14 (70) 3 (50) 25 (68)
      Female 2 (40) 1 (25) 6 (30) 3 (50) 12 (32)
      Not reported 9 7 9 31 7 63
      Pet 11 (79) 8 (89) 5 (38) 22 (50) 5 (42) 51 (55)
      Stray 3 (21) 1 (11) 8 (62) 22 {50) 7 (58) 4 (45)
      Not reported 7 1 8
      Unlicensed 10 4 4 19 5 42
      Not reported 4 5 9 32 8 58
      Neutered 3 (43) 3 (21)
      Not neutered 2 (100) 1 (100) 2 (100) 4 (57) 2 (100) 11 (79)
      Not reported 12 11 44 11 86
      Positive Rabies
      Exposure (FRA) 3 (7) 3 (4)
      Negative Rabies
      Exposure (FRA) 9 (100) - 4 (100) 10 (100) 38 (93) 11 (100) 72 (96)
      Not reported 5 5 3 10 2 25
      ” Circumstances of Attack
      Unprovoked 11 (79) 1 (11) 7 (15) 19 (22)
      Climbed on person 1 (9) 1 (2) 2 {2}
      Handling/petting 1 (7) 6 (67) 5 (45) 21 (46) 4 {67} 37 (43)
      Feeding 1 (9) 4 (9) 5 (6)
      Finger in cage 1 (7) 1 (9) 2 (2)
      Otherwise provoked 1 (7) 2 (22) 3 (27) 13 (28) 2 (33) 21 (24)
      Not reported 2 5 7 14

      “ fay have included tetanus prophylaxis.

      Nee”

      -47-
      Year

      1978
      1979
      1980
      1981
      1982
      1983
      1984
      1985
      1986
      1987

      Total

      NUMBER OF FERRET ATTACKS REPORTED
      IN RESPONSE TO CDHS REQUEST FOR DATA IN EARLY

      Table 4

      Reporting State

      California

      3

      3
      14
      23
      31
      _26

      100

      Oregon

      20

      13
      10
      22

      76

      * Reporting from two counties only
      ** Data had been lost by Oregon health officials although

      48

      kk

      kk

      1986

      Arizona*

      17
      53
      51
      34
      49
      37

      249

      cases were reported to have occurred in these years.
      Table 5

      Reported Ages of Ferret Attack Victims

      Age of

      Victim California Oregon Arizona Totals*
      "Baby" 5 5
      <1yr. 6 4 5 15
      1-2 yrs. 4 5 \
      >2-3 yrs. 4 \ 8 2 32
      "Child" Z

      >3-4 yrs. J 2 P.
      >4-5 yrs. 3 7 1 7
      >5-6 yrs. 2 3

      >6-7 yrs. ] 2

      >7-8 yrs. +64 ; +342
      >8-9 yrs. 200**

      >9-10 yrs. 3

      "Adult" 63 J J
      Unknown 14 17 31
      Total 100 76 249 425

      * 13 percent of attacks were in children 4 years old or younger.

      ** Presumably in this age range.

      49
      Table 6

      Estimated Annual Ferret Attack Rates Per Million Persons

      California Oregon
      Estimated attacks/year 27 20
      (from Table 4)
      1986 State population 27 Quek
      (millions)
      Attacks/million persons ] 7.4
      (Persons)

      * Reporting from two counties only

      50

      Arizona*

      50

      25
      ~<
      c
      [e*)
      3

      1958
      1978

      198]
      1982
      1982

      1983
      1985

      1985

      1986

      1986

      1986

      1987

      Table 7

      Rabies-Infected Pet European Ferrets Reported in the

      State

      Kentucky

      South Carolina

      North Dakota
      Kansas

      Virginia

      Wisconsin

      California

      Michigan

      South Carolina

      Washington, D.C.

      North Dakota

      Towa

      United States

      Details/Circumstances

      None available

      Child bitten in face by
      ferret from pet shop

      Pets; no vaccination
      Pet; no vaccination

      Ferret fought with

      raccoon in yard

      None available

      Developed rabies after
      recapture; had escaped
      3 months earlier after
      imported from Colorado.

      (Possible error in
      diagnosis) FRA-positive
      but adult mouse test
      negative; live virus

      vaccine-induced?

      Bit owner who “spanked” it;
      purchased in a North Carolina
      pet shop 1 month earlier;
      Unneutered 5 month old male

      "Kissing" exposure of person;
      Purchased from pet shop

      1 year earlier

      Developed rabies after re-
      capture; Never vaccinated

      Ferret maintained in a cage
      as a breeder; was bred

      earlier in year

      51

      Reference

      CDC 1983
      Diesch 1981

      CDC 1983
      CDC 1983
      Jenkins 1985

      CDC 1985
      VPHU 1986

      Anderson 1986

      CDC 1986a

      CDC 1986a

      CDC 1987

      Currier 1987
      The authors gratefully acknowledge help from the following persons in

      ACKNOWLEDGMENTS

      compiling data on which this report is based:

      M.
      G.
      E.
      B.
      A.
      Ss.
      L.
      S.
      M.
      M.
      P.
      V.
      W.
      D.
      L.


      mrAw

      .

      WMmRaoONragan

      K. Abelseth
      R. Anderson

      F. Baker, Jr.

      Benda
      Chandler
      Coffey
      Curtis

      L. Diesch
      Eidson
      Ford
      Fisher

      A. Giles
      H. Griffith
      I. Herbet
      Hunter
      Klein
      Kwan

      S. Mahoney
      Miller
      Mock

      - Pape

      Porter
      E. Reynolds

      - Ruprecht

      Sawyer
      Shumway

      - Tacal

      K. Weeks

      . Wise

      -52-

      . Alishouse

      Avedian
      M. Beck
      Bunter
      A. Clark
      Cookson
      Davis

      . M. Doll
      - Elliott
      . Fishbein

      Freeman

      A. Greco,
      K. Grigor
      T. Horman
      S. Kelly
      Kurth
      Liska
      Maier
      Minor
      Murrill
      Pearson
      Purves

      J. Ruff
      Rutty
      Scott

      E. Sorhage
      Walsh

      P. Williams
      E. Wright
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      Veterinary Public Health Unit. 1986. Rabid ferret, Tehama County,
      California. California Department of Health Services memo to the

      record. February 7, 1986.

      Volobuev, V. T., D. Ternovsky and A. Graphodatsky. 1974. Taxonomic

      status of the ferret (Putorius putorius furo) by karyological data.

      Zool. Zh. 53:1738-1740.

      Weeks, E. K. 1986. County of Los Angeles Department of Health

      Services reports to Veterinary Public Health Unit, CDHS. April 1986.

      Wellstead, G. 1982. Ferrets and Ferreting. T.F.H. Publications,

      Neptune City, New Jersey.

      Wiktor, T. J. and H. Koprowski. 1978. Monoclonal antibodies against
      rabies virus produced by somatic cell hybridization: Detection of

      antigenic variants. Proc. Natl. Acad. Sci. USA, 75: 3938-3942.

      Williams, L. P. 1986a. Oregon State Health Department reports to
      Veterinary Public Health Unit, CDHS. January 31, February 14 and 20,

      1986.

      Williams, L. P. 1986b. Oregon State Health Department warns of
      danger of keeping pet ferrets in home with infants. News Release,

      Department of Human Resources. September 17, 1986.

      Willis, L. S. and M. V. Barrow. 1971. The ferret (Mustela putorius

      furo L.). Lab. Anim. Sci.: 21:712-716.

      -64-
      113.

      114.

      115.

      Winsted, Vv. 1981. Ferrets. T.F.H. Publications, Neptune City,

      New Jersey.

      World Health Organization Expert Committee on Rabies. 1984. Seventh

      Report. World Health Organization, Tech. Rep. Ser. 709.

      Wright, M. E. 1986. Arizona Department of Health Services reports on

      ferret bites sent to Veterinary Public Health Unit, CDHS, in

      March 1986.

      -65-
      APPENDIX I

      CALIFORNIA STATE L

      RELEVANT TO FERRETS

      66
      § 2116 FISH AND GAME GENERALLY
      Article 1
      GENERALLY
      Section

      2116. Wild animal defined.

      2116.5. Legislative findings and intent.

      2117. Enforcing officer defined. 7

      2118. Importation, transportation, possession and release of specified wild
      animals; permit.

      2118.2. Elk; importation; findings.

      9118.3. Elk horns or antlers; removal from live elk for commercial purposes.

      2118.4, Elk; importation; seizure.

      2118.5. Possession without permit.

      2119. Publication of list. ; ; ;

      2120. Regulations governing entry, transportation, keeping, etc., of restricted
      wild animals; possession of other wild animals. —

      212), Freeing or permitting escape or release of restricted wild animals.

      2122, Regulations in co-operation with department of food and agriculture.

      2123. Descriptive material concerning restricted wild animals.

      Historical Note

      For general discussion of prior game
      laws, see Historical Note preceding § 3000.

      Cross References

      Exclusion of specified species from definition of “aquaculture”, regulation under this
      chapter, see § 17.

      § 2116. Wild animal defined

      As used in this chapter, “wild animal” means any animal of the class
      Aves (birds), class Mammalia (mammals), class Amphibia (frogs, toads,
      salamanders), class Osteichtyes (bony fishes), class Monorhina (lam-
      preys), class Reptilia (repitles), class Crustacea (crayfish), or class Gas-
      tropoda (slugs, snails) which is not normally domesticated in this state as
      determined by the commission.

      (Stats.1957, c. 456, p. 1348, § 2116. Amended by Stats.1961, c. 617, p. 1770, § 2;
      Stats.1974, c. 15038, p. 3296, 8 1.)

      Legislative Counsel Notes

      “Wild animal” used in place of “wild bird
      or animal.”

      Historical Note

      The 1961 amendment included the classes The 1974 amendment substituted “is not
      Amphibia, Osteichtyes, Monorhina, Reptilia normally domesticated in this state for ei-
      and Gastropoda; and deleted “phylum Mol- ther is not normally domesticated in this
      lusca (snails)’”’. State or not normally native to this State”.

      236

      IMPORTATION AND TRANSPORTATION § 2117
      th. 2
      Similar to § 1165 of the Fish and Game Section 1165 was derived from Stats.1933
      Code of 1933, added by Stats.1953, ¢. 178, p.e, 76 87. ae
      1105, § 1. y ¢ p c. 76, p. 516, & 7.
      Forms

      See West’s California Code Forms, Fish and Game.

      Cross References

      Importation of wild animals by permit, application to excluded animals under this chapter,
      see Health and Safety Code § 259948.

      § 2116.5. Legislative findings and intent

      The Legislature finds and declares that wild animals are being cap-
      tured for importation and resale in California; that some populations of
      wild animals are being depleted; that many animals die in captivity or
      transit; that some keepers of wild animals lack sufficient knowledge or
      facilities for the proper care of wild animals; that some wild animals are
      a threat to the native wildlife or agricultural interests of this state; and
      that some wild animals are a threat to public health and safety. It is the
      intention of the Legislature that the importation, transportation, and
      possession of wild animals shall be regulated to protect the health and
      welfare of wild animals captured, imported, transported, or possessed, to
      reduce the depletion of wildlife populations, to protect the native wildlife
      and agricultural interests of this state against damage from the exist-
      ence at large of certain wild animals, and to protect the public health and
      safety in this state.

      (Added by Stats.1974, c. 1503, p. 3296, § 1.5.)

      Library References

      Game @3'/..
      C.J.S. Game § 7.

      Notes of Decisions

      1. In general raccoons for the pet trade. 65 Ops.Atty.
      The fish and game commission has the Gen. 648, 12-30-82.
      authority to prohibit the captive breeding of

      § 2117. Enforcing officer defined

      As used in this chapter, “enforcing officers’? means the enforcement
      personnel of the department, the state plant quarantine officers, and the
      county agricultural commissioners.

      (Stats.1957, c. 456, p. 1343, § 2117.)

      Legislative Counsel Notes

      No change.

      237
      § 2117 FISH AND GAME GENERALLY
      Div. 3

      Historical Note
      Identical with § 1165.1 of the Fish and Section 1165.1 was derived from Stats.

      Game Code of 1938, added by Stats.1953, ¢. 1933, ¢. 76, p. 517, § 8.
      178, p. 1105, § 1.

      Cross References

      Deputies and county fish and game wardens, see § 850 et seq.

      § 2118. Importation, transportation, possession and release of spe-
      cified wild animals; permit
      It is unlawful to import, transport, possess, or release alive into this
      state, except under a revocable, nontransferable permit as provided in
      this chapter and the regulations pertaining thereto, any wild animal of
      the following species:
      (a) Class Aves: (birds)
      Family Cuculidae (cuckoos)
      All species.
      Family Alaudidae (larks)
      Skylark, Alauda arvensis
      Family Corvidae (crows, jays, magpies)
      All species.
      Family Turdidae (thrushes)
      European blackbird, Turdus merula
      Missel (or mistle), thrush, Turdus viscivorus
      Family Sturnidae (starlings and mynas or mynahs)
      All species of the family, except hill myna (or hill mynah)
      Gracula religiosa (sometimes referred to as Eulabes religiosa)
      Family Ploceidae (weavers)
      The following species:
      Spanish sparrow, Passer hispaniolensis
      Italian sparrow, Passer italiae
      European tree sparrow, Passer montanus
      Cape sparrow, Passer capensis
      Madagascar weaver, Foudia madagascariensis
      Baya weaver, Ploceus baya
      Hawaiian rice bird, Munia nisoria
      Red-billed quelea, Quelea quelea
      Red-headed quelea, Quelea erythrops
      Family Fringillidae (sparrows, finches, buntings)
      Yellowhammer, Emberiza citrinella
      (b) Class Mammalia (mammals)
      Order Primates
      All species except those in family Homonidae
      Order Edentata (sloths, anteaters, armadillos, etc.)
      All species,
      238

      IMPORTATION AND TRANSPORTATION § 2118
      Ch. 2

      Order Marsupialia (marsupials or pouched mammals)
      All species.
      Order Insectivora (shrews, moles, hedgehogs, etc.)
      All species.
      Order Dermoptera (gliding lemurs)
      All species.
      Order Chiroptera (bats)
      All species.
      Order Monotremata (spiny anteaters, platypuses)
      All species.
      Order Pholidota (pangolins, scaly anteaters)
      All species.
      Order Lagomorpha (pikas, rabbits, hares)
      All species, except domesticated races of rabbits.
      Order Rodentia (rodents)

      All species, except domesticated golden hamsters, also known as
      Syrian hamster, Mesocricetus auratus; domesticated races of rats or
      mice (white or albino; trained, dancing or spinning, laboratory-reared);
      and domestic strains of guinea pig (Cavia porcellus).

      Order Carnivora (carnivores)

      All species, except domestic dogs (Canis familiaris) and domestic
      cats (Felis catus).

      Order Tubulidentata (aardvarks)

      All species.

      Order Proboscidea (elephants)

      All species.

      Order Hyracoidea (hyraxes)

      All species.

      Order Sirenia (dugongs, manatees)

      All species.

      Order Perissodactyla (horses, zebras, tapirs, rhinoceroses, etc.)

      All species except those of the family Equidae.

      Order Artiodactyla (swine, peccaries, camels, deer, elk, except elk
      (genus Cervus) which are subject to Section 2118.2, moose,
      antelopes, cattle, goats, sheep, etc.)

      All species except: domestic swine of the family Suidae; American
      bison, and domestic cattle, sheep and goats of the family
      Bovidae; races of big-horned sheep (Ovis canadensis) now or
      formerly indigenous to this state.

      Mammals of the orders Primates, Edentata, Dermoptera, Monotrema-
      ta, Pholidota, Tubulidentata, Proboscidea, Perissodactyla, Hyracoidea,
      Sirenia and Carnivora are restricted for the welfare of the animals,
      except animals of the families Viverridae and Mustelidae in the order
      Carnivora are restricted because such animals are undesirable and a

      239
      § 2118 FISH AND GAME GENERALLY
      Div. 3
      menace to native wildlife, the agricultural interests of the state, or to the
      public health or safety.
      (c) Class Amphibia (frogs, toads, salamanders)
      Family Bufonidae (toads)
      Giant toad or marine toad, Bufo marinus
      (d) Class Monorhina (lampreys)
      All species.
      (e) Class Osteichthyes (bony fishes)
      Family Serranidae (bass)
      White perch, Morone or Roccus americana
      Family Clupeidae (herring)
      Gizzard shad, Dorosoma cepedianum
      Family Sciaenidae (croakers)
      Freshwater sheepshead, Aplodinotus grunniens
      Family Characidae (characins)
      Banded tetra, Astyanax fasciatus
      All species of piranhas
      Family Lepisosteidae (gars)
      All species.
      Family Amiidae (bowfins)
      All species.
      (f) Class Reptilia (snakes, lizards, turtles, alligators)
      Family Crocodilidae
      All species.
      (g) Class Crustacea (crustaceans)
      Genus Cambarus (crayfishes)
      All species.
      Genus Astacus (crayfishes)
      All species.
      Genus Astacopsis (crayfishes)
      All species.
      (h) Class Gastropoda (slugs, snails, clams)
      All species of slugs.
      All species of land snails.

      (i) Such other classes, orders, families, genera, and species of wild
      animals which may be designated by the commission in cooperation with
      the Department of Food and Agriculture, (a) when such class, order,
      family, genus or species is proved undesirable and a menace to native
      wildlife or the agricultural interests of the state, or (b) to provide for the
      welfare of wild animals.

      (j) Classes, families, genera, and species in addition to those listed
      above may be added to or deleted from the above lists from time to time

      240

      IMPORTATION AND TRANSPORTATION § 2118
      Ch. 2

      by commission regulations in cooperation with the Department of Food
      and Agriculture.

      (Stats.1957, ¢. 456, p. 1343, § 2118. Amended by Stats.1961, c. 617, p. 1770, § 3;
      Stats.1970, c. 302, p. 578, § 1; Stats.1974, c. 1503, p. 3296, § 2; Stats.1977, c. 436,

      p. 1470, § 1; Stats.1979, c. 1074, p. 3844, § 1, eff. Sept. 28, 1979.)

      Legislative Counsel Notes

      “Wild animal” used in place of ‘wild bird
      or animal.”

      Historical Note

      The 1961 amendment rewrote the section,
      which prior thereto read:

      “It is unlawful to import or transport
      alive into this State, except as provided in
      this chapter, any wild animal of any of the
      following species or groups:

      “(a) Yellowhammer, Emberiza citrinella;
      hooded crew, Corvus cornix; carrion crow,
      Corvus corone; rook, Corvus frugilegus;
      European starling, Sturnus vulgaris; crest-
      ed starling, Aethiopsar cristatellus; com-
      mon mynah, Acridotheres tristis; skylark,
      Alauda arvensis; European blackbird, Tur-
      dus merula; missel thrush, Turdus viscivo-
      rus; baya, Ploceus baya; Madagascar
      weaver, Foudia madagascariensis.

      “(b) Opossums of the family Didelphidae;
      European rabbit, Lepus cuniculus; Europe-
      an hare, Lepus europaeus, and all other
      species of the family Leporidae except
      domesticated races of rabbits; bank vole,
      Clethrionomys glareolus; field voles, Micro-
      tus hirtus and Microtus agrestis; water rat,
      Arvicola amphibus; long-tailed field mouse,
      Apodemus sylvaticus; and aJ] other species
      of the following rodent families, Muridae
      (mice and rats), Cricetidae (hamsters and
      old world field mice), Sciuridae (ground
      squirrels, prairie dogs, woodchucks, etc.),
      and Geomyidae (pocket gophers); weasel,
      Mustela nivalis; stoat, Mustela erminea;
      ferret, Mustela furio; mongoose, Herpestes
      mungo, and all other species of the genus
      Herpestes; European mole, Talpa europaea,
      and all other Talpidae; flying foxes or fruit
      bats of the family Pteropodidae; bats of the
      family Desmodontidae.

      “(c) Crayfishes of the genera Cambarus
      and Astacus; slugs and land snails of the
      mollusean class Gastropoda.

      ‘(d) Such other species of wild animal
      which may be subsequently designated by
      the commission when such species are

      proved undesirable and a menace to the
      native wildlife or to the agricultural inter-
      ests of this State.”

      The 1970 amendment made “possession”
      of the listed species unlawful.

      The 1974 amendment inserted “revocable,
      nontransferable” preceding ‘‘permit” in the
      introductory sentence; rewrote subd. (b),
      which prior thereto read:

      “Class Mammalia (mammals)
      “Family Didelphidae (opossums)
      “All species
      “Family Leporidae (rabbits, hares)

      “All species, except domesticated
      races of rabbits

      “Family Cricetidae (hamsters, field
      mice, voles)

      “All species, except laboratory-reared
      golden hamsters, also known as
      Syrian hamster, Mesocricetus aura-
      tus

      “Family Muridae (mice, rats)

      “All species, except domesticated
      races (white or albino) of rats and
      mice

      “Family Geomyidae (pocket gophers)

      “All species

      “Family Sciuridae (squirrels, wood-
      chucks)

      “All species

      “Family Dasyproctidae (agoutis)
      “AIL species

      “Family Procyonidae (raccoons, coatis)
      “All species

      “Family Mustelidae (weasels, ferrets)
      “All species

      “Family Viverridae (civets, mongooses)
      “All species

      241
      § 2118

      “Family Talpidae (moles)

      FISH AND GAME GENERALLY
      Div. 3

      The 1977 amendment, in subd. (b), under
      “Order Rodentia”, substituted “‘domesticat-

      ii All species : ‘ ed” for “laboratory-reared’’ preceding
      Family Pteropodidae (flying foxes or “golden hamsters”, and deleted “nutria pos-
      fruit bats) sessed in accordance with provisions of the
      “All species Food and Agricultural Code” following
      “Family Desmodontidae (true vampire “Mesocricetus auratus”.
      bats) The 1979 amendment, in subd. (b), under
      “All species “Order Artiodactyla”, inserted “except elk

      “Family Cervidae (deer, elk, moose)
      “All species

      “Pamily Bovidae (cattle, buffaloes, bi-
      son, sheep, goats, gazelles, Old
      World and African antelopes)

      “All species, except domesticated cat-
      tle, sheep, goats, and the races of
      big-horned sheep (Ovis canadensis)
      now or formerly indigenous to this

      (genus Cervus) which are subject to Section
      2118.2”.

      Section 4.5 of Stats.1979, c. 1074, p. 3845,
      provides:

      “Any person prior to January 1, 1980,
      may apply to the State Board of Control
      for, and the board shall pay, compensation
      for actual damages, which shall not include
      any anticipated loss of profits, not to exceed

      “ State ‘ . one hundred fifty thousand dollars
      Family Tayassuidae (peccaries) ($150,000), as a result of establishing a busi-
      “All species ness in reliance on a permit issued by the

      “Family Suidae (swine)
      “All species, except domesticated
      swine”;

      substituted ‘Department of Food and Agri-
      culture” for “State Department of Agricul-
      ture” and “Department of Agriculture” in
      subds. (i) and (j), respectively; inserted des-
      ignation (a) within subd. (i); and added, to
      the end of subd. (i), “or (b) to provide for
      the welfare of wild animals”.

      Department of Fish and Game, pursuant to
      approval granted by the Fish and Game
      Commission, when it is no longer legal to
      conduct such business.”

      Similar to § 1165.2 of the Fish and Game
      Code of 1933, added by Stats.1953, c. 178, p.
      1105, § 1.

      Section 1165.2 was derived from Stats.
      1933, c. 76, p. 515, § 1.

      Cross References

      Inspection and refusal of admittance of restricted wild animals, see § 2185. .
      Permits for importation or transportation of certain wild animals designated by this

      section, see § 2150.

      Supply of descriptive material concerning restricted wild animals, see § 2123.

      Administrative Code References
      Permits for research purposes, see 14 Cal.Adm. Code 671.1 et seq.

      Library References

      Fish 8.
      Game 3'/.

      CJ.5. Fish § 26.
      C.J.S. Game § 7.

      Notes of Decisions

      In general 2

      1. Validity

      IMPORTATION AND TRANSPORTATION § 2118.4

      Ch. 2

      piranha which he was ordered to destroy or
      otherwise dispose of, and as he thus ac-
      quired no protectible property right in the
      fish, the order and the legislation and regu-
      lations pursuant to which the order was
      issued did not deprive appellant of his prop-
      erty without due process of law. Adams v.
      Shannon (1970) 86 Cal.Rptr. 641, 7 C.A.3d
      427.

      2. In general

      In suit wherein tropical fish dealer
      sought an injunction to restrain the depart-
      ment of fish and game from enforcing stat-
      utes and regulations prohibiting the impor-

      tation and possession of piranha, the record
      established that legislature’s action was
      based upon a firm foundation in fact, there
      being qualified expert opinion that piranha,
      if introduced into the waters of California,
      could seriously endanger existing species of
      aquatic life. Adams v. Shannon (1970) 86
      Cal.Rptr. 641, 7 C.A.3d 427.

      3. Raccoons

      The fish and game commission has the
      authority to prohibit the captive breeding of
      raccoons for the pet trade. 65 Ops.Atty.
      Gen. 648, 12-30-82.

      § 2118.2. Elk; importation; findings

      Except as provided in Section 1007, it is unlawful to import any elk
      (genus Cervus) into this state. The department may import elk pursuant
      to Section 1007, if prior to such importation, the department issues
      written findings justifying the need for and explaining the purpose of the

      importation.

      This section shall not apply to zoos certified by the United States

      Department of Agriculture.

      (Added by Stats.1979, ¢. 1074, p. 3844, § 2, eff. Sept. 28, 1979.)

      Library References

      Game 3'2.
      C.J.S. Game § 7.

      § 2118.3. Elk horns or antlers; removal from live elk for commer-

      cial purposes

      No part of any elk horn or antler shall be removed from any live elk

      for commercial purposes.

      (Added by Stats.1979, c. 1074, p. 3845, § 3, eff. Sept. 28, 1979.)

      Library References

      Game 3'.
      C.J.S. Game § 7.

      § 2118.4. Elk; importation; seizure
      The department shall seize any elk imported in violation of Section

      2118.2.

      (Added by Stats.1979, c. 1074, p. 3845, § 4, eff. Sept. 28, 1979.)

      Raccoons 3 As appellant, a tropical fish dealer, im-
      Validity 1 ported in violation of California law the 140

      242

      243
      § 671 FISH AND GAME COMMISSION TITLE 14
      (p. 52.72.4) (Register 87, No. 19—6-0-87)

      (h) Any permit issued pursuant to these peigulations may be cancelled or
      sospendal at any time by the director of the department when, in his judg-
      ment, permittee is acting or has acted contrary to the terms and conditions of
      subject permit, or if, in his judgment, the safety or welfare of the species
      authorized to be taken by subject permit is or may be jeopardized by the actions
      of permittee.
      NOTE: Authority cited: Section 1002, Fish and Game Code. Reference: Sections 3511,
      4700, 5050 and 5515, Fish and Game Code.
      HISTORY:

      1. New section filed 2-18-81; effective thirtieth day thereafter (Register 81, No. 8).

      671. Importation, Transportation and Possession of Wild Animals.

      The following species of the families which are prohibited or for which a
      permit is required are determined to be not normally domesticated in this state
      and shall not be imported into, transported within or possessed in this state, and
      permits for their entry, transportation or possession will be refused by the
      Department of Fish and Game, except that permits may be granted for the
      species as specified herein and for purposes designated below subject to the
      conditions and restrictions contained in Sections 671.1, 671.2, 671.4, 671.5, and/or
      671.6 and such other conditions as may be designated by the department.

      The birds, amphibians, fish, reptiles, crustaceans, and gastropods which are
      prohibited or for which a permit is required in this regulation are restricted to
      reduce the depletion of these wild animal populations, or because such animals
      are undesirable and a menace to native wildlife, the agricultural interests of the
      state, or to the public health or safety.

      Mammals of the orders Primates, Edentata, Dermoptera, Monotremata,
      Pholidota, Tubulidentata, Proboscidea, Perissodactyla, Hyracoidea, Sierenia,
      and Carnivora, which are prohibited or for which a permit is required are
      restricted for the welfare otf the animals, except ani of the families Viver-
      ridae, Procyonidae except pandas and Procyon lotor (American raccoon), and
      Mustelidae excepting genera Amblonyx, Aonyx, Pteronura and Lutra (river
      otters), and the family Felidae in the order Carnivora are restricted because
      such animals are undesirable and a menace to native wildlife, the agricultural
      interests of the state, or to the public health or safety. Mammals of other orders
      which are prohibited or for which a permit is required except the families
      Macropodidae (kangaroos, wallabies) and Phalangeridae (Phalanger, koalas)
      in the order Marsupialia, and the family Camelidae (camels, etc.) in the order
      Artiodactyla are restricted because auch animals are undesirable and a menace
      to native wildlife, the agricultural interests of the state, or to the public health.

      Subject to applicable provisions of the Fish and Game Code or regulations of
      the Fish and Game Commission set forth in Title 14, species not listed below
      may be imported, ale hea or possessed without a permit. No person shall
      release into the wilds of this state any animal which is not native to California
      except as provided in these regulations. (See Section 671.5.)

      (a) Class Aves—Birds

      (1) Family Alaudidae—Larks

      Alauda arvensis (Skylark) —Prohibited.

      (2) Family Cuculidae—Cuckoos

      All species—Prohibited.

      @) Family Corvidae—Crows, Ravens, Rooks, Jackdaws (Genus Corvus)

      All species—Prohibited. .
      TITLE 14 FISH AND GAME COMMISSION § 671
      {Register 67, No. 19—6-0-87) (p. 52.72.5)

      (4) Family Turdidae—Thrushes, Blackbirds, Fieldfare
      (A) Turdus merula (European blackbird) —Prohibited.
      (B) Turdus viscivorus ae thrush) —Prohibited.
      (C) Turdus pilaris (Fieldfare)—Prohibited.
      (D) Turdus musicus (Song thrush)—Prohibited.
      & Family Sturnidae—Starlings, Mynahs
      species except Sturnus is (Starling) and Gracula religiosa or
      Eulabes religiosa (Hill mynahs)—Prohibited.
      (6) F “mil y Ploceidae—Sparrow, Weavers, Queleas
      (A) Genus Passer (Sparrow)
      All species except Passer domesticus (English house sparrow) -—Prombites,
      (B) Foudia madagascariensis (Madagascar weaver)—Prohibited.
      (C) Ploceus baya (Baya weaver)—Prohibited.
      (D) Genus Quelea (Quelea)—All species prohibited.
      (7) Family Estrildidae—Waxbills, Munias, Ricebirds
      (A) Padda oryzivora (Java sparrow)—Prohibited.
      (B) Munia nisoria (Hawaiian rice bird)—Prohibited.
      (8) Family Emberizidae—Yellowhammer
      Emberiza citrinella (Yellowhammer)—Prohibited.
      (9) Order Falconiformes—Falcons, Eagles, Hawks, Vultures
      ies prohibited, except under conditions set forth in Section 670, Title
      14, California Administrative Code. .
      (10) Order Stigfones—Owis
      Alpes prohibited, except under conditions set forth in Section 670, Title
      14, California Administrative le.
      (

      11) Family Pyenonotidae—Bulbuls or Fruit Thrushes
      Pycnonotus jocosus (Red-whiskered bulbul)—Prohibited.
      (12) Family Zosteropidae—Whiteeyes
      Genus Zosterops—All species prohibited.

      (13) Family Psittacidae—Parrots, Parakeets

      Myiopsitta monachus (Monk or Quaker parakeet) —Prohibited.

      (b) Class Mammalia—Mammals

      (1) Order Primates
      species except Family Hominidae—Prohibited.

      @ Order Edentata—Sloths, anteaters, armadillos, etc.
      species—Prohibited.

      @ Order Marsupialia—Marsupials or Pouched Animals
      species except Didelphis marsupialis (Common opossum)—Prohibited.

      (4) Order Insectivora—Shrews, Moles, Hedgehogs, etc.
      species—Prohibited.

      & Order Dermoptera—Gliding Lemurs
      species—Prohibited.

      & Order Chiroptera—Bats
      species—Prohibited.

      @ Order Monotremata—Spiny Anteaters, Platypuses
      Species—Prohibited.

      @ rder Pholidota—Pangolins, Scaly Anteaters
      species—Prohibited.
      § 671 FISH AND GAME COMMISSION TITLE 14
      (p. 52.72.6) (Register 87, No. 19—6-0-87)

      @) Order Lagomorpha—Pikas, Rabbits, and Hares
      species except domesticated races of rabbits and hares of the Family
      Leporidae—Prohibited.

      (10) Order Rodentia—Hamster, Field Mice, Voles, Muskrats, Gerbils, Squir-
      rels, Chipmunks, Woodchucks, and Prairie Dogs

      (A) species protibited except:

      1. Ondatrazibethica (Muskrats)—Under conditions set forth in Fish and
      Game Code Section 2250;

      2. Myocastor coypus (Nutria)—Under conditions set forth in Food and Agri-
      cultural Code Sections 165-165.6;

      3. Domesticated races of golden hamsters of the species Mesocricetus aura-
      tus;

      4. Domesticated races of rats or mice (white or albino; trained, dancing or
      spinning, laboratory-reared); and

      5. Domesticated races of guinea pigs of the species Cavia porcellus.

      6. Domesticated races of chinchillas of the species Chinchilla laniger.

      (11) Order Carnivora—Raccoons, Ringtailed Cats, Kinkajous, Coatis, Caco-
      mistles, Weasels, Ferrets, Skunks, Polecats, Stoats, Mongoose, Civets, Wolves,
      Foxes, Coyotes, Lions, Tigers, Ocelots, Bobcats, Servals, Leopards, Jaguars,
      Cheetahs, Bears, etc.

      (A) Skunks (all species) —Prohibited except under conditions set forth in
      Section 2606.8, Title 17, California Administrative Code.

      (B) Family Felidae—All species except Felis catus (domestic cats) —Prohib-
      ited except as provided in Sections 671.1 and 671.4(b) of these regulations and
      Section 3005.9, Fish and Game Code, except that permits are required for
      cheetahs (Acinonyx).

      (C) All other species except Canis familiaris (domestic dogs) —Prohibited.

      (12) Order Tubulidentata—Aardvarks

      All species—Prohibited.

      (13) Order Proboscidae (Elephants)

      All species—Prohibited.

      (14) Order Hyracoidae (Hyraxes)

      All species—Prohibited.

      (15) Order Sirenia (Dugongs, Manatees)

      All species—Prohibited.

      (16) Order Perissodactyla (Horses, Zebras, Tapirs, Rhinoceroses, etc.)

      All species except Family Equidae—Prohibited.

      (17) Order Artiodactyla—Swine, Peccaries, Camels, Deer, Elk, Moose, An-
      telopes, Cattle, Goats, Sheep, etc.

      (A) Family Cervidae

      Elk (Cervus)—Prohibited, except a permit may be issued to a California
      licensed domesticated game breeder for possession of domesticated elk already
      within California.

      All other species—Prohibited, except permits may be issued for all species to
      a California licensed domesticated game breeder.

      (B) Family Bovidae

      All species—Prohibited, except permits may be issued to a California licensed
      domesticated game breeder for races of Ovis canadensis (Bighorn sheep)
      which are now or were formerly indigenous to this state.
      TITLE 14 FISH AND GAME COMMISSION , § 671
      (Register 88, No. 92-27-68) (p. 52.72.7)

      (C) Family Antilocapriade—Pronghorn Antelope Prohibited, except per-
      mits may be issued to a California licensed domesticated game breeder.
      (D) All other families and species except Bison bison (American bison) and
      domesticated swine, cattle, sheep, or goats—Prohibited.
      (c) Class Amphibia—Frogs, Toads, Salamanders
      (1) Family Bufonidae—Toads
      Bufo marinus, Bufo paracnemis, Bufo horribilis (Giant toad or marine toad
      group) and all other large toads from Mexico and Central and South America—
      Prohibited.
      @ Family Pipidae—Tongueless Toads
      species of genus Xenopus—Prohibited.
      (d) Class Agnatha—Jawless Fishes
      (1) Family Petromyzontidae—Lampreys
      nonnative species—Prohibited.
      (e) Class Osteichthyes—Bony Fishes
      (1) Family Bescienthritee— emperate Basses
      Morone americana (White perch)—Prohibited.
      (2) Family Clupeidae—Herrings
      Dorosoma cepedianum (Gizzard shad)—Prohibited.
      (3) Family Sciaenidae—Drums
      Aplodinotus Ejunniens (freshwater drum)—Prohibited.
      (4) Family Characidae—Characins
      (A) Astyanax fasciatus (Banded tetra) —~Prohibited.
      (B) All species of genera Serrasalmus, Serrasalmo, Pygocentrus, Taddyella,
      Rooseveltiella, and Pygopristis (Piranhas)—Prohibited.
      (C) Hoplias malabaricus (Tiger fish)—Prohibited.
      5) Family Lepisosteidae—Gars
      species—Prohibited.
      6) Family Amiidae—Bowfins
      species~—Prohibited.
      (1) Family Poeciliidae—Livebearers
      Belonesox belizanus (Pike top minnow)—Prohibited.
      i Family (Channidae)—Snakeheads
      : s en hima ”
      (9) ily rinidae—Carps or Minnows
      (A) Leuciscus idus or—Prohibited.
      (B) Ctenopharyngodon idellus (Grass carp) —Prohibited.
      (10) Family Trichomycteridae (Pygidiidae)—Parasitic Catfishes.
      All species—Prohibited.
      (11) Family Cetopsidae—Whalelike Caifishes.
      All species—Prohibited.
      (12) Family Clariidae—Airbreathing Catfishes
      a species of genera Clarias, Dinotopterus, and Heterobranchus —Prohibit-

      (13) Family Heteropneustidae (Saccobranchidae)—Airsac Catfishes

      All species—Prohibited.

      (14) Family Cichlidae—Cichlids

      (A) Tilapia sparrmanii (Tilapia) —Prohibited.

      (B) Tilapia zillii (Redbelly tilapia)—Prohibited, except permit may be is-
      sued to a person or agency for importation, transportation, or possession in the
      counties of San Bernardino, Los Angeles, Orange, Riverside, San Diego, and
      mperial.
      § 671 FISH AND GAME COMMISSION TITLE 14
      (p. 52.72.8) (Register 88, No. 92-27-48)

      (15) Family Anguillidae—Freshwater Eels.

      All appeits of genus Anguilla—Prohibited.

      (f) Class Chondrichthyes—Cartilaginous Fishes

      (1) Family Carcharhinidae—Requiem sharks

      All species of genus Carcharhinus (Freshwater sharks) —Prohibited.

      (2) Family Potamotrygonidae—River stingrays

      All species Prohibited’

      (g) Class Reptilia—Reptiles

      (1) Order Crocodilia—Crocodiles, Caimans, Alligators and Gavials

      All species—Prohibited.

      (2) Family. Chelyridae—Snapping turtles

      All species—Prohibited.

      (3) Family Elapidae—Cobras, Coral Snakes

      All species—Prohibited.

      (4) amily Viperidae—Adders and Vipers

      All species—Prohibited.

      (5) Family Crotalidae—Pit Vipers .

      All species—prohibited, except Crotalus viridis (western rattlesnake) , Crota-
      Jus atrox (western diamondback rattlesnake), Crotalus ruber (red diamond
      rattlesnake), Crotalus scutulatus (Mojave rattlesnake), Crotalus mitchellis
      (speckled rattlesnake) and Crotalus cerastes (sidewinder)

      (6) Family Colubridae—Colubrids

      (A) Dispholidus typus (Boomslang)—Prohibited.

      (B) Theoltornis kirtlandi (Bird or Vine Snake) —Prohibited.

      (h) Class Crustacea—Crustaceans

      (1) All species of family Cambaridae except Procambarus clarkii and Or-
      conectes virilis—Prohibited.

      (2) All species of genus Eriocheir—Prohibited.

      (i) Class Gastropoda—Slugs, Snails

      All nonnative species of slugs and land snails except Rumina decollata (decol-
      late snail) in the counties of San Bernardino, Riverside, Imperial, Orange, San
      Diego, Los Angeles, Ventura, Tulare and Santa Barbara with the concurrence
      of the appropriate county agricultural commissioners; and Helix aspersa
      (brown garden snail)—Prohibited.

      NOTE: Authority cited: Sections 1002, 2116, 2118, 2120, 2122, 3005.9 and 3005.92, Fish and
      Game Code. Reference: Sections 1002, 2116-2118, 2118.2, 2118.4, 2119-2155, 2185-2191,
      3005.9 and 3005.92, Fish and Game Code.

      HISTORY:

      1. Amendment of subsection (g) (5) filed 2-9-84; effective thirtieth day thereafter
      (Register 84, No. 6). For prior history, see Register 81, No. 29.

      2. Amendment of subsection (i) filed 7-8-85; effective thirtieth day thereafter (Register
      85, No. 28).

      3. Editorial correction of NOTE filed 9-20-85; effective thirtieth day thereafter (Regis-
      ter 85, No. 38).

      4. Notice of Erroreous Filing filed 3-24-86 by OAL; purported amendment of subsec-
      tion (g) (5) filed in error on 2-5-86 is null and void and text as filed with Secretary of State
      on 2-9-84 remains in effect uninterrupted (Register 86, No. 13).

      5. Amendment of subsection (g) (5) filed 3-24-86; effective thirtieth day thereafter
      (Register 86, No. 13).

      6. Amendment of subsection (h) filed 11-7-86; effective upen filing (Register 86,
      No. 43).

      7. Amendment of subsection (g) (1) filed 5-1-87; operative 5-31-87 (Register 87, No. 19).

      8. New subsection (e) (15) filed 2-16-88; operative 3-17-88 (Register 88, No. 9).
      TITLE 14 FISH AND GAME COMMISSION § 671.2
      (Register 88, No. 92-27-88) (p. 52.72.9)

      671.1. Permits for Zoological Gardens, Research and Film Making.

      (a) With the concurrence of the Departments of Health and Food and
      Agriculture, the Department of Fish and Game may issue permits for importa-
      tion, transportation and possession of restricted and prohibited species listed
      herein for exhibition by zoological gardens and for use for scientific or public
      health research by a college, university, or government research agency, or
      other bona fide scientific institution as determined by the department, to meet
      immediate research or medical needs. With the above concurrence the depart-
      ment may issue permits authorizing the importation, transportation, and
      possession of restricted or prohibited species listed herein if such animals are
      to be used for public display or public exhibition through the mediums of
      motion pictures and television; no permittee shall transfer possession or owner-
      ship of any such animal without approval of the department. All animals import-
      ed and/or possessed under this subsection must be imported, transported,
      maintained, and disposed of under such conditions as the department may
      prescribe,

      NOTE: Authority cited: Sections 1002, 2116, 2118, 2118.5, 2120, 2122, 3005.9 and 3005.92,
      Fish and Game Code. Reference: Sections 2002, 2116-2118, 2118.2, 2118.4, 2119-2155,
      2185-2191, 3005.9 and 3005.92, Fish and Game Code. :

      HISTORY: ° .

      i. Amendment filed 5-6-75 as an emergency; effective upon filing (Register 75, No. 18).
      For prior history, see Register 75, No. 4.

      2. Certificate of Compliance filed 7-3-75 (Register 75, No. 27).

      3. Amendment of subsection (a) filed 5-11-79; effective thirtieth day thereafter (Regis-
      ter 79, No. 19).

      4. Amendment of NOTE filed 7-16-81; effective thirtieth day thereafter (Register 81,
      No. 29).

      5. Order of Repeal of subsection (b) filed 6-3-85 by OAL pursuant to Government Code
      Section 11349.7; effective thirtieth day thereafter (Register 85, No. 26).

      6. Editorial correction of NOTE filed 9-20-85; effective thirtieth day thereafter (Regis-
      ter 85, No. 38).

      671.2. Neutered Male Animals.

      NOTE: Authority cited: Sections 1002, 2116, 2118, 2120, 2122, 3005.9 and 3005.92, Fish and
      Game Code. Reference: Sections 1002, 2116-2118, 2118.2, 2118.4, 2119-2155, 2185-2191,
      3005.9 and 3005.92, Fish and Game Code.

      HISTORY:
      1. Amendment filed 6-30-66; effective thirtieth day thereafter (Register 66, No. 20).
      2. Amendment filed 1-24-75 as an emergency; effective upon filing. Certificate of
      Compliance included (Register 75, No. 4).
      3. New NOTE filed 7-16-81; effective thirtieth day thereafter (Register 81, No. 29).
      4, Editorial correction of NOTE filed 9-20-85; effective thirtieth day thereafter (Regis-
      ter 85, No. 38}.
      5. Repealer filed 7-15-87; operative 8-14-87 (Register 87, No. 29).

    Attachments

    • Constantine and Kizer - 1988 - Pet European ferrets a hazard to public health, small livestock and wildlife.pdf

      Report

    • Snapshot

    Related

    • Pet ferrets--a hazard to public health and wildlife.
  • Pet ferrets--a hazard to public health and wildlife.

    Item Type Journal Article
    Author K W Kizer
    Author D G Constantine
    Date 1989-4
    Library Catalog PubMed Central
    URL https://www.ncbi.nlm.nih.gov/pmc/articles/PMC1026598/
    Accessed 7/14/2023, 10:51:10 AM
    Extra PMID: 2735056 PMCID: PMC1026598
    Volume 150
    Pages 466
    Publication Western Journal of Medicine
    Issue 4
    Journal Abbr West J Med
    ISSN 0093-0415
    Date Added 7/14/2023, 10:51:12 AM
    Modified 5/22/2025, 8:01:59 PM

    Attachments

    • Kizer and Constantine - 1989 - Pet ferrets--a hazard to public health and wildlife..pdf
    • PubMed Central Link
    • Snapshot

    Related

    • Pet European ferrets: a hazard to public health, small livestock and wildlife
  • Pleistocene Refugia and Holocene Expansion of a Grassland-Dependent Species, the Black-Footed Ferret ( <i>Mustela nigripes</i> )

    Item Type Journal Article
    Author Samantha M. Wisely
    Author Mark J. Statham
    Author Robert C. Fleischer
    Date 2008-02-19
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://academic.oup.com/jmammal/article-lookup/doi/10.1644/07-MAMM-A-077.1
    Accessed 5/21/2025, 4:32:20 PM
    Volume 89
    Pages 87-96
    Publication Journal of Mammalogy
    DOI 10.1644/07-MAMM-A-077.1
    Issue 1
    Journal Abbr J Mammal
    ISSN 0022-2372, 1545-1542
    Date Added 2/6/2014, 2:04:05 PM
    Modified 5/21/2025, 8:44:10 PM

    Attachments

    • Wisely et al. - 2008 - Pleistocene Refugia and Holocene Expansion of a Grassland-Dependent Species, the Black-Footed Ferret.pdf
  • Potential risks associated with the legalization of exotic predators such as the ferret (Mustela putorius furo) in California

    Item Type Conference Paper
    Author Thomas, G. Moore
    Author Desley, A. Whisson
    Editor R. O. Baker
    Editor A. C. Crabb
    Abstract The interest in possessing ferrets as pets has given rise to controversy between the "rights" of the individual to own the pet of their choice and the concerns for protection of wildlife in California. An overview of the legislative history in California illustrates the state's attempts at protecting native wildlife species from exotic wild birds and animals. Concerns as to the potential threats associated with the legalization of ferrets in California are warranted in light of the wildlife damage resulting from the deliberate introduction of ferrets in New Zealand and the non-native red fox in California. A framework to assess risks involved with introducing non-native species that may impact native wildlife is needed.
    Date 1998
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://escholarship.org/uc/item/4fh248bk
    Accessed 4/29/2019, 10:53:39 AM
    Volume 18
    Place Davis, CA
    Pages 96-99
    Proceedings Title Proceedings of the Vertebrate Pest Conference
    Conference Name 18th Vertebrate Pest Conference
    DOI 10.5070/V418110186
    Date Added 4/29/2019, 10:53:39 AM
    Modified 2/13/2023, 8:31:25 AM

    Attachments

    • Moore and Whisson - 1998 - Potential risks associated with the legalization o.pdf
  • Reassortment between avian H5N1 and human H3N2 influenza viruses in ferrets: a public health risk assessment

    Item Type Journal Article
    Author Sara Jackson
    Author Neal Van Hoeven
    Author Li-Mei Chen
    Author Taronna R. Maines
    Author Nancy J. Cox
    Author Jacqueline M. Katz
    Author Ruben O. Donis
    Date 2009
    Short Title Reassortment between avian H5N1 and human H3N2 influenza viruses in ferrets
    Library Catalog Google Scholar
    Extra Publisher: Am Soc Microbiol
    Volume 83
    Pages 8131–8140
    Publication Journal of virology
    Issue 16
    Date Added 7/14/2023, 12:17:57 PM
    Modified 10/22/2025, 7:58:58 AM

    Attachments

    • Full Text
    • Full Text PDF
  • Separating Wild from Domestic American Mink <i>Neovison vison</i> Based on Skull Morphometries

    Item Type Journal Article
    Author Ashley L. Tamlin
    Author Jeff Bowman
    Author David F. Hackett
    Date 09/2009
    Language en
    Library Catalog CrossRef
    URL http://www.bioone.org/doi/abs/10.2981/08-004
    Accessed 8/4/2014, 9:02:51 AM
    Volume 15
    Pages 266-277
    Publication Wildlife Biology
    DOI 10.2981/08-004
    Issue 3
    ISSN 0909-6396
    Date Added 8/4/2014, 9:02:51 AM
    Modified 6/14/2025, 9:26:21 AM

    Attachments

    • Tamlin et al. - 2009 - Separating Wild from Domestic American Mink iNeo.pdf
  • Some behavioural differences between the European polecat, Mustela putorius, the ferret, M. furo, and their hybrids

    Item Type Journal Article
    Author Trevor B. Poole
    Abstract The aim of this work was to identify behavioural differences between the domesticated ferret Mustela furo and its wild counterpart the European polecat M. putorius and to obtain objective measures of one or more of these differences. The animals used in this study were tame M. putorius, M. furo, wild caught hybrids and laboratory reared F1 hybrids between the two species. Ferrets differed in their behaviour in unfamiliar environments and in the fact that they do not develop fear of man. The F1 hybrids resembled M. putorius in that they develop a fear of man if left with their mother during a critical period between 7½ and 8½ weeks of age. The phenomenon of imprinting may be involved in this developmental process. The attention response to a rustling noise was investigated under controlled conditions revealing differences in response decrement between the four types of polecat. M. putorius and the hybrids habituated more rapidly to the sound than did the ferrets. The results obtained for F1 hybrids depended upon the previous environmental history of the individual. Those individuals which had been living in outdoor cages responded differently from the ones in indoor cages. The hypothesis was put forward that the hybrids respond more frequently if the experimental situation provides a greater contrast with their normal environment and that the individual's response threshold is therefore “set” to specific environmental conditions. The results obtained agree with the hypothesis first put forward by Lorenz (1953) that the behaviour of domesticated animals resembles that of juvenile individuals of their wild counterpart. Summary: A comparison is made between the behaviour of European polecats (Mustela putorius), the domesticated ferret (M. furo) and hybrids between the two species. Differences were found in exploratory behaviour, fear of man and the frequency of occurrence of the “attention response”. The occurrence of the “attention response” was studied under controlled conditions. It was found, on playing the rustling sound 40 times at minute intervals, that this sound elicited the attention response for longer in ferrets than in European polecats and hybrids between the two species. The most rapid habituation occurred in polecats, the F1 hybrids being intermediate in their response between M. putorius and M. furo. The results obtained for the F1 hybrids were influenced by the conditions under which the animals were previously caged. The results appear to support Lorenz' view that domesticated animals show juvenility in their behaviour as compared with their wild counterparts.
    Date 1972
    Language en
    Library Catalog Wiley Online Library
    URL https://onlinelibrary.wiley.com/doi/abs/10.1111/j.1469-7998.1972.tb04073.x
    Accessed 2/20/2025, 6:02:37 PM
    Extra _eprint: https://onlinelibrary.wiley.com/doi/pdf/10.1111/j.1469-7998.1972.tb04073.x
    Volume 166
    Pages 25-35
    Publication Journal of Zoology
    DOI 10.1111/j.1469-7998.1972.tb04073.x
    Issue 1
    ISSN 1469-7998
    Date Added 2/20/2025, 6:02:37 PM
    Modified 10/22/2025, 11:30:12 AM

    Attachments

    • Full Text PDF
    • Poole - 1972 - Some behavioural differences between the European polecat, Mustela putorius, the ferret, M. furo, an.pdf
    • Snapshot
  • Space use, resource selection and territoriality of black-footed ferrets: implications for reserve design

    Item Type Journal Article
    Author David A. Eads
    Author Dean E. Biggins
    Author Travis M. Livieri
    Author Joshua J. Millspaugh
    Date 03/2014
    Language en
    Short Title Space use, resource selection and territoriality of black-footed ferrets
    Library Catalog CrossRef
    URL http://www.bioone.org/doi/abs/10.2981/wlb.13070
    Accessed 6/24/2014, 9:48:04 AM
    Volume 20
    Pages 27-36
    Publication Wildlife Biology
    DOI 10.2981/wlb.13070
    Issue 1
    ISSN 0909-6396, 1903-220X
    Date Added 6/24/2014, 9:48:04 AM
    Modified 5/21/2025, 8:43:21 PM

    Attachments

    • Eads et al. - 2014 - Space use, resource selection and territoriality o.pdf
  • Spatial patterns of co-occurrence of the European wildcat Felis silvestris silvestris and domestic cats Felis silvestris catus in the Bavarian Forest National Park

    Item Type Journal Article
    Author Tanja Beutel
    Author Björn Reineking
    Author Annika Tiesmeyer
    Author Carsten Nowak
    Author Marco Heurich
    Abstract After persecution and a long period of extinction in parts of central Europe, the European wildcat is currently increasing its range, also to areas deemed unsuitable for this species. This range expansion facilitates close contact with domestic cats, which can reach high population densities even in protected areas. We provide unambiguous evidence that the European wildcat is present in the Bavarian Forest National Park. We assessed the frequency of domestic cat occurrence, and analyzed the distributions of the two felid subspecies with regard to residential areas, forested habitat, elevation and protected areas, and analyzed their spatial overlap. Camera traps installed in the national park detected six putative wildcats in 2008–2015 at elevations between 800 and 1100 m a.s.l. Genetic analysis of material obtained from hair traps with valerian-treated lure sticks confirmed the presence of three wildcat individuals in early 2015. The number of wildcat events detected increased slightly in recent years and wildcats were detected closer to shrub cover and the forest edge. Of the domestic cat events, 90% were within 1.1 km of residential areas, but some moved up to nearly 3 km into the national park. Ranges of wildcats and domestic cats broadly overlapped. All but one camera trap that recorded wildcats also recorded domestic cats, and some camera traps recorded domestic cats but no wildcats. Domestic cats were the fifth most often detected mammal species in the protected area. To avoid a negative impact of domestic cats on wildcats through hybridization, which might already occur, and considering the ecological impact of predation by domestic cats, we recommend a buffer zone of 1 km surrounding the national park, where domestic cats should not be allowed outdoors.
    Date January 1, 2017
    Library Catalog bioone.org (Atypon)
    URL http://www.bioone.org/doi/abs/10.2981/wlb.00284
    Accessed 3/30/2017, 12:24:33 PM
    Pages wlb.00284
    Publication Wildlife Biology
    DOI 10.2981/wlb.00284
    Issue 2017
    Journal Abbr Wildlife Biology
    ISSN 0909-6396
    Date Added 3/30/2017, 12:24:33 PM
    Modified 6/14/2025, 9:31:19 AM

    Attachments

    • Beutel et al. - 2017 - Spatial patterns of co-occurrence of the European .pdf
    • Snapshot
  • Summer/autumn movements, mortality rates and density of feral ferrets (Mustela furo) at a farmland site in North Canterbury, New Zealand

    Item Type Journal Article
    Author Peter Caley
    Author Grant Morriss
    Abstract [For two summer/autumn periods (1999, 2000), we studied the movements and survival of feral ferrets (Mustela furo L.) at a site in North Canterbury that had been previously subjected to intensive control of ferrets. Movement distances of juvenile ferrets from the place of initial to final capture were generally low (median =1.2 km) though variable [$\bar{X}$=2.5±1.0 (±S.E.M.), range 0.1–21.7 km]. The estimated instantaneous mortality rate of juvenile ferrets was high ($\bar{X}$ = 0.8 yr−1), though imprecise (95% C.I. 0.4–1.8 yr−1), as was that of adult ferrets ($\bar{X}$=1.1 yr−1, 95% C.I. 0.3–4.5yr−1). Summer/autumn home ranges of adult ferrets in North Canterbury were larger than estimates from comparable studies. The density of ferrets during May was 1.7 km−2 in the first year and 2.5 km−2 in the second year. This corresponded with a trend for juvenile movements to increase, and juvenile survival rates to decrease. Our results suggest that formal studies to determine whether movements and survival of juvenile ferrets are density-dependent are warranted.]
    Date 2001
    Archive JSTOR
    Library Catalog JSTOR
    URL https://www.jstor.org/stable/24056362
    Accessed 4/29/2019, 10:48:03 AM
    Volume 25
    Pages 53-60
    Publication New Zealand Journal of Ecology
    Issue 1
    ISSN 0110-6465
    Date Added 4/29/2019, 10:48:03 AM
    Modified 5/21/2025, 8:43:10 PM

    Attachments

    • Caley and Morriss - 2001 - Summerautumn movements, mortality rates and densi.pdf
  • The chronology of a sad historical misjudgement: The introductions of rabbits and ferrets in nineteenth-century New Zealand

    Item Type Journal Article
    Author Carolyn King
    Abstract During the second half of the nineteenth century, rabbit damage to pastoral runs was threatening to bring about a serious economic crisis in the southern South Island of New Zealand. The obvious solution was to import thousands of the rabbits’ ‘natural enemies’—starting with ferrets—in the hope they would keep the rabbits under control. This article reviews the arguments surrounding this decision, and describes the persons most actively involved, their reasoning and their actions, compiled from contemporary documents and reports, many unpublished. The introduction of ferrets proceeded despite repeated local and international warnings, but ferrets could not remove enough rabbits to prevent the continuing damage to sheep pastures. Contemporary ecologists often wonder why such reckless decisions were made, but this new analysis of the historic details illustrates how well‑meant and apparently reasonable environmental management actions can precipitate a cascade of unintended and undesirable consequences.
    Date 2017-06-23
    Language en
    Short Title The chronology of a sad historical misjudgement
    Library Catalog DOI.org (Crossref)
    URL http://press-files.anu.edu.au/downloads/press/n2519/pdf/08_king.pdf
    Accessed 5/21/2025, 4:31:20 PM
    Volume 3
    Pages 139-173
    Publication International Review of Environmental History
    DOI 10.22459/IREH.03.01.2017.09
    Issue 1
    Journal Abbr IREH
    ISSN 22053204, 22053212
    Date Added 7/14/2023, 12:20:16 PM
    Modified 5/24/2025, 7:06:14 AM

    Attachments

    • King - 2017 - The chronology of a sad historical misjudgement The introductions of rabbits and ferrets in ninetee.pdf
  • The Cranial Morphology of the Black-Footed Ferret: A Comparison of Wild and Captive Specimens

    Item Type Journal Article
    Author Tyler Antonelli
    Author Carissa L. Leischner
    Author Adam Hartstone-Rose
    Abstract The black-footed ferret (Mustela nigripes), a North American mustelid species, was once found abundantly throughout the Midwest until the extreme decline in prairie dogs (Cynomys spp.), the black-footed ferret’s primary food source, brought the species to near-extinction. Subsequently, the Black-Footed Ferret Recovery Program was created in the 1980s with a goal of bringing all remaining individuals of the species into captivity in order to breed the species back to a sustainable population level for successful reintroduction into the wild. While many components of the ferrets’ health were accounted for while in captivity—especially those affecting fecundity—this study aims to assess the effects that captivity may have had on their cranial morphology, something that has not been widely studied in the species. In a previous study, we showed that the captive ferrets had significant oral health problems, and here we aim to document how the captive diet also affected their skull shape. For this study, 23 cranial measurements were taken on the skulls of 271 adult black-footed ferrets and 53 specimens of two closely related species. Skulls were divided based on sex, species, captivity status and phase of captivity and compared for all measurements using stepwise discriminant analysis as well as principal component analysis derived from the combined variables. We found that there are significant differences between captive and wild specimens, some of which are larger than interspecific variation, and that a diet change in the captive specimens likely helped decrease some of these differences. The results suggest that captivity can cause unnatural cranial development and that diet likely has a major impact on cranial morphology.
    Date 2022-10-09
    Language en
    Short Title The Cranial Morphology of the Black-Footed Ferret
    Library Catalog DOI.org (Crossref)
    URL https://www.mdpi.com/2076-2615/12/19/2708
    Accessed 5/21/2025, 4:32:28 PM
    Rights https://creativecommons.org/licenses/by/4.0/
    Volume 12
    Pages 2708
    Publication Animals
    DOI 10.3390/ani12192708
    Issue 19
    Journal Abbr Animals
    ISSN 2076-2615
    Date Added 5/21/2025, 4:32:28 PM
    Modified 5/21/2025, 4:32:28 PM

    Attachments

    • Antonelli et al. - 2022 - The Cranial Morphology of the Black-Footed Ferret A Comparison of Wild and Captive Specimens.pdf
  • The current distribution and potential for future range expansion of feral ferret Mustela putorius furo in Ireland

    Item Type Journal Article
    Author Daniel J. Buckley
    Author Mathieu Lundy
    Abstract Predicting the potential invaded range of a non-native species is an important tool for conservation biology. The ferret (Mustela putorius furo) is a known invasive species outside its native range and has recently been confirmed as a feral species in Ireland. To determine the current distribution of feral populations, an all-island survey was conducted during 2006-2008. Using the results of this survey, a landscape modelling approach, using presence-only data was applied to predict the potential future range of this species in Ireland, given the availability of suitable habitat. The results suggest that Ireland appears to be potentially highly suitable for ferrets and, therefore, the possible ecological impacts of this species in Ireland are discussed.[PUBLICATION ABSTRACT]
    Date Jun 2013
    Language English
    Library Catalog ProQuest
    URL https://search.proquest.com/docview/1355115940/abstract/B02A8DE21D54546PQ/1
    Accessed 4/29/2019, 10:50:45 AM
    Rights Springer-Verlag Berlin Heidelberg 2013
    Volume 59
    Pages 323-330
    Publication European Journal of Wildlife Research; Heidelberg
    DOI http://dx.doi.org/10.1007/s10344-012-0677-4
    Issue 3
    Journal Abbr Eur J Wildl Res
    ISSN 16124642
    Date Added 4/29/2019, 10:50:45 AM
    Modified 5/26/2025, 10:33:15 PM

    Tags:

    • Animal populations
    • Landscape ecology
    • Dispersal
    • Mammals
    • Animal Welfare
    • Nonnative species
    • Sports And Games

    Attachments

    • Buckley and Lundy - 2013 - The current distribution and potential for future .pdf
  • The diet of an invasive nonnative predator, the feral ferret , and implications for the conservation of ground-nesting birds

    Item Type Journal Article
    Author Thomas W. Bodey
    Author Stuart Bearhop
    Author Robbie A. McDonald
    Abstract Introduced carnivores have had a significant impact on the fauna of a number of countries, particularly on islands. In the British Isles, several offshore islands holding internationally important aggregations of seabirds and shorebirds support self-sustaining feral ferret populations, often as the top terrestrial predator. However, little is known about the interactions between ferrets and both native and nonnative prey in these locations. We examined the diet of feral ferrets on Rathlin Island, Northern Ireland. We determined the frequency of occurrence of prey items and constructed energetic models to determine their potential impact on both native and introduced prey. Rabbits occurred in 75% of scats, while birds, carrion, and brown rats were important secondary items. There was little difference between the diets of males and females. Estimates of the energy requirements of the population at current, and with hypothetically reduced, rabbit availability revealed the potential for carrion to maintain the ferret population over winter. Management options could thus focus on reducing anthropogenic food sources as an immediate way of mitigating the threat to ground-nesting birds, while other strategies, including eradication, are considered.
    Date June 2010
    Library Catalog HAL Archives Ouvertes
    URL https://hal.archives-ouvertes.fr/hal-00599783
    Accessed 4/29/2019, 10:49:30 AM
    Rights http://www.springer.com/tdm
    Volume 57
    Pages 107-117
    Publication European Journal of Wildlife Research
    DOI 10.1007/s10344-010-0404-y
    Issue 1
    Journal Abbr Eur J Wildl Res
    ISSN 1612-4642, 1439-0574
    Date Added 4/29/2019, 10:49:30 AM
    Modified 5/26/2025, 10:32:44 PM

    Tags:

    • Invasive species
    • Management
    • Hyperpredation
    • Island
    • Mustelid

    Attachments

    • Bodey et al. - 2010 - The diet of an invasive nonnative predator, the fe.pdf
  • The ferret and its diseases

    Item Type Journal Article
    Author L. M. Ryland
    Author J. R. Gorham
    Date 1978-11-01
    Language en
    Library Catalog PubMed
    Extra PMID: 367994
    Volume 173
    Pages 1154-1158
    Publication Journal of the American Veterinary Medical Association
    Issue 9
    Journal Abbr J Am Vet Med Assoc
    ISSN 0003-1488
    Date Added 5/22/2025, 8:08:11 PM
    Modified 5/22/2025, 8:10:06 PM

    Tags:

    • Animals
    • Carnivora
    • Male
    • Tuberculosis
    • Pregnancy
    • Diet
    • Dogs
    • Ferrets
    • Female
    • Reproduction
    • Alopecia
    • Botulism
    • Congenital Abnormalities
    • Copulation
    • Distemper
    • Enteritis
    • Estrus
    • Housing, Animal
    • Orthomyxoviridae Infections
    • Rabies
    • Scabies
    • Staphylococcal Infections
    • Streptococcal Infections

    Notes:

    • The Ferret and Its Diseases

      Lennox M. Ryland, BS, and John R. Gorham, DVM, PhD

      THE FERRET (Mustela putorius furo), a carnivore of the
      family Mustelidae, was domesticated from the wild Eu-
      ropean polecat and first imported into the United States
      around 1875. Its history can be traced back as far as
      the 4th century BC, when it was first used in Europe
      for exterminating rodents and snakes and later for hunt-
      ing rabbits. Two varieties of ferrets are recognized,
      based on coloration: the wild type, or fitch ferret, is
      pale yellow-buff with black mask, limbs, and tail; the
      albino ferret is white with pink eyes, although an in-
      tact male may appear yellow as a result of profuse seba-
      ceous secretion. The albino is genetically recessive to
      the wild phenotype.

      The fitch ferret is not to be confused with the
      American black-footed ferret (Mustela nigripes), which
      once inhabited the prairies throughout much of the
      United States. Now on the list of rare and endangered
      species published by the Department of the Interior,}
      its near extinction may be at least partially explained
      by decreased numbers of its principal food, the prairie
      dog. Today, only 4 black-footed ferrets are known to
      remain, and these are in captivity.

      Both wild type and albino varieties have been used
      as laboratory research animals, in which capacity they
      have contributed to studies of the physiology of repro-
      duction and of the immunology, pathology, and phar-
      macology of viral diseases, virus-induced neoplasms, and
      bacterial infections. Their ease of maintenance, small
      size, and generally friendly disposition make them
      equally suitable as household pets, especially if they can
      be raised from infancy in close human contact.

      Species Characteristics

      In our ferret colony at the Experimental Fur Farm
      at Washington State University, adult ferrets range from
      600 to 2,000 g in weight and 44 to 56 cm in length (nose
      to tip of tail). Males frequently are nearly twice as
      large as females. Large (830%-40%) seasonal fluctua-
      tions in weight are due to the accumulation of subcu-
      taneous fat in the autumn and its subsequent loss in
      the spring and summer. Pelt cycling is concurrent with
      photoperiodic weight variation, but there is no obvious
      color change between winter and summer coats. Ferrets
      normally live for 5 to 6 years, although life spans of 8
      to 9 years are common. The young are born hairless
      and blind, their eyes begin to open at 3 to 4 weeks,
      and they reach their adult weight at 4 months.

      From the Pioneering Research Laboratory, Science and Education Admin-
      istration, US Department of Agriculture, Pullman, WA 99164.

      Mention of a trademark or proprietary product does not constitute a guar-
      antee or warranty of the product by the US Department of Agriculture and does
      not imply its approval to the exclusion of other products that also may be
      suitable.

      » Google

      As do all members of the Mustelidae, ferrets have
      paired musk-producing glands at the base of the tail,
      which secrete when the animal is angry or excited and,
      in females, at estrus. These can be removed by a vet-
      erinarian at the time of neutering if the ferret is to be
      a pet, although some musky odor from skin secretions
      will persist. An unusual anatomic variation is a single
      central artery (innominate artery I) that arises from
      the descending part of the aortic arch and proceeds
      along the midline ventral to the trachea through the
      neck. Here it divides into the left carotid artery and
      innominate artery II, which then gives rise to the right
      subclavian and carotid arteries. The postulation is that
      this midline vessel serves to prevent occlusion of cerebral
      blood flow when the ferret turns its head 180 degrees.?

      Male ferrets are called “hobs” and females “‘jills.’’
      Unlike their feral polecat relatives, they are not strictly
      nocturnal animals and do not develop an inherent fear
      of man. Although their eyes are not adapted for color
      vision, they have been trained in laboratories to dis-
      criminate between objects, and their performance is re-
      ported to be similar to that of primates.®

      Reproduction

      Ferrets reach sexual maturity in the spring after
      their birth, or at approximately 1 year of age. The fe-
      male ferret is scasonally polyestrous, with a normal
      breeding season from March to August. When ferrets
      are maintained on a natural photoperiod, the onset of
      estrus is triggered by increasing day length, but both on-~
      set and duration of estrus can be modified by manipula-
      tion of photoperiod. For instance, 1st estrus may be ac-
      celerated by exposing immature females to long periods
      of light after they have been reared on short days; some
      immature males may also be brought into breeding con-
      dition of a shorter duration by similar treatment.* Par-
      adoxically, too much light may be inhibitory for both
      sexes. Male ferrets typically begin and end their po-
      tential breeding seasons earlier than the female (De-
      cember to July), a disparity which may be a functional
      adaptation to allow time for sperm maturation.

      Onset of estrus in the female is easily recognized by
      enlargement of the vulva, which swells to full size within
      a month and regresses to normal size within 2 to 8 weeks
      after coitus. The female will remain in estrus up to
      120 days if not bred. Estrus can be artificially termin-
      nated after 10 days by parenteral injection of 1,000 usp
      units of chorionic gonadotropin’; if vulvar swelling does
      not subside within 1 week, the dose may be repeated.”

      8 Western Medical Supply, Inc, Arcadia, Calif.
      >» Marshall G, North Rose, NY: Personal communication, 1978.

      JAVMA, Vol 173, No. 9
      Coitus is prolonged and may take from 1 to 3 hours.
      Ovulation is induced 30 to 35 hours after coitus, and
      the sperm may penetrate the ovum while it is still in
      the follicle. Unlike other Mustelidae, implantation is
      not delayed; the trophoblast attaches to the uterine epi-
      thelium on the 12th or 13th day after coitus. The ges-
      tation period is 42 days (pseudopregnancy of 40 to 42
      days if fertilization fails), and the female will return to
      estrus after the young are weaned 6 to 8 weeks after
      delivery, thereby allowing production of 2 litters an-
      nually. Mean litter size is 8 (range 2 to 17), but the
      fertility of both males and females will decline after 3
      to 4 years of age.®

      Housing

      In the laboratory or breeding establishment, ferrets
      may be housed outdoors, or if breeding is to be artifi-
      cially controlled, indoors where exposure to light can
      be regulated. They may be kept singly or in sexually
      distinct groups, although males may become belligerent
      when housed together, especially during the breeding
      season. Females in estrus should not be housed together
      if breeding is desired, because their play may induce
      ovulation and pseudopregnancy. In our colony, most
      ferrets are housed singly unless they are to be bred;
      then, the female is transferred to the male’s cage and
      left only long enough for the animals to copulate. This
      is to prevent the male from excessively grabbing the
      female’s neck and thus inflicting wounds which may be-
      come secondarily infected. Double-breeding the female
      with more than 1 male on the same day or on 2 con-
      secutive days results in maximal fertility. The females
      are then removed and are later placed in larger cages
      equipped with nesting boxes to. ensure privacy at par-
      turition and during lactation.

      Cages may be contructed from heavy wire mesh,
      which will permit most waste materials to fall into col-
      lecting troughs or pans below, or they may have solid
      flooring covered with wood shavings or newspaper. The
      mesh will be more easily cleaned and disinfected. The
      cage should be provided with an enclosed area in which
      the ferret can sleep undisturbed. A deep plastic box in
      which a hole has been cut to afford entry makes a sat-
      isfactory den and may be suspended from the roof or
      wall of the cage. Because one of the corners of the cage
      will habitually be used for droppings, a pet ferret may
      be easily trained to a litter box.

      Ferrets have been shown to thrive at temperatures
      varying from near freezing to 32C.° Because adults
      do not have well-developed sweat glands and are prone
      to heat exhaustion, as manifested in panting and flaccid
      inactivity, hot and humid conditions should be min-
      imized by adequate circulation of air. Outdoor pens
      must be shaded from the sun, and when temperatures
      approa’ .g 22C are common, a sprinkler system for
      the roof is advisable. Although bedding is otherwise
      inessential, wood shavings, excelsior, sugarcane pulp, or
      othe» insulatory material should be provided when tem-
      peratures are near freezing, and the pregnant females
      should be given similar materials 2 weeks before parturi-
      tion. Special exercise equipment or runs need not be
      provided, although pet ferrets will benfit from regular,

      November I, 1978

      Google

      supervised freedom and have even been trained to walk
      outdoors with a cat harness and leash.

      Diet

      The ferret is a carnivore and may be satisfactorily
      maintained on mixtures of fish, chicken, turkey, horse-
      meat, animal by-products, cereal, and vitamin supple-
      ment. If a ferret is to be fed large amounts of fish,
      which contain unsaturated fatty acids, or oxidized horse
      fat, its diet should be supplemented with stabilized vi-
      tamin E so that it receives at least 10 mg of vitamin E
      per day. Pet ferrets may thrive on canned cat or dog
      food or dry cat or dog food fed ad libitum to prevent
      irritability associated with hunger. Cod liver oil (38 to
      5 drops/day) may be added to the diet should the coat
      become dry. Milk and bread are also palatable additions
      to the ration, and milk is a good source of supplemental
      calcium, although it may cause loose feces. The ferret
      has little, if any, capacity to digest fiber. Restricted
      quantities of table scraps may be offered to pets, but
      small bones can become lodged in their gastrointestinal
      tracts and should be withheld from the diet. Water, in
      either a cup or drinking bottle, should be available at
      all times and changed daily.

      Routine Veterinary Procedures

      Ferrets must be routinely protected against canine
      distemper by intramuscular or subcutaneous vaccination
      with modified live virus of chicken embryo tissue culture
      origin. Vaccination every other year is sufficient after
      a primary vaccination at 10 weeks of age (4 weeks of
      age if kits are from susceptible, unvaccinated dams).
      Vaccine containing a high titer of modified live distem-
      per virus will protect a ferret almost immediately. Be-
      cause distemper is almost invariably fatal to ferrets and
      because many dogs are carriers of the disease, the ferret
      should not be brought into a clinic until after it is
      vaccinated. Killed tissue vaccine provides only short-
      term immunity that is slow to develop and is not effec-
      tive in all ferrets.‘ Vaccine prepared from ferret cell
      culture must not be used because the attenuated virus
      may retain its virulence for its natural host.

      Rabies vaccine, killed only and of murine origin,
      should be administered annually, beginning at 3 months.
      Ferrets raised commercially should be inoculated at
      weaning with type C botulism toxoid, which will pro-
      tect against type C botulism for about 1 year. Pet
      ferrets need not be vaccinated if their food is kept fresh
      to prevent growth of Clostridium botulinum. Because
      ferrets are not susceptible to feline panleukopenia, ca-
      nine hepatitis, or mink virus enteritis, they do not re-
      quire vaccination against these diseases.

      Intramuscular injections should be given, as with
      a dog or cat. The ferret is best restrained when it is
      grasped firmly around its neck, shoulders, and forelimbs.
      Some people prefer to pick up their own ferrets with
      bare hands, although commercial breeders often use a
      light pair of leather gloves, especially if the animal is
      nervous or excited.

      Pet males should be castrated to reduce aggressive-

      © United, Animal Science Division, Mogul Corp, Middleton, Wis, or Amer-
      ican Scientific Laboratories, Madison, Wis.

      1155
      ness and desire to roam, and females should be spayed
      at 6 to 8 months of age. Scent glands can be removed
      at the same time; standard procedures for anal gland
      removal should be followed. Ether (0.25 ml/kg), so-
      dium pentobarbital (35 mg/kg, intraperitoneally), and
      ketamine HCI (9 parts:1 acetylpromazine, and given at
      the rate of 35 mg/kg, 1m) are suitable anesthetics. Ket-
      amine is routinely used in our laboratory for procedures
      requiring anesthesia of 30 minutes or less. The ferret’s
      normal respiratory rate is 33 to 36 breaths/minute and
      rectal temperature varies between 37.8 and 40 C. Blood
      may be obtained by toenail clipping (1 to 2 ml), jugular
      venipuncture, or cardiac tap. Packed cell volume is 35%
      to 40%, total red blood cell count is 6.5 to 8.5 x 10°/
      mm*, and white cell count is 6 to 9 x 10°/mm?.§

      Diseases

      Viral Diseases—Ferrets are highly susceptible to ca-
      nine distemper. Moreover, because the case fatality rate
      approaches 100%, ferrets are used as test animals for
      detection of the virus. After exposure, the ferret appears
      normal until the 7th, 8th, or 9th day, when it loses its
      appetite, eating little or nothing from then on. The
      eyelids stick together, and the eyes and nose exude a
      mucopurulent discharge on the 8th, 9th, or 10th day.
      A rash may appear beneath the chin and in the inguinal
      region on the 10th, 11th, or 12th day after exposure,
      and the foot pads may swell and become hyperkeratotic.
      Commonly, the anus protrudes. The animal’s condition
      continues to deteriorate until death at 12 to 14 days
      after exposure with ferret-adapted strains (3 to 3%
      weeks with canine strains). Ferrets that survive the
      catarrhal phase may die in a neurotropic episode, signs
      of which include hyperexcitability, excess salivation,
      muscular tremor, convulsions, and coma. Because the
      signs of distemper are so typical, the disease is rarely
      confused with anything else. However, clinical diagnosis
      can be confirmed by virus isolation or by examination
      for characteristic inclusion bodies in the urinary bladder
      and trachea.

      Ferrets are also susceptible to infection with several
      strains of human influenza virus, which may cause initial
      signs similar to those of distemper. Within 48 hours
      of exposure, the affected ferret becomes quiet and list-
      less, its rectal temperature rises sharply to 40 or 41 C,
      its nose becomes moist, and it refuses food. Fever lasts
      for 1 day, then returns on the 3rd day. Sneezing attacks
      may develop, accompanied by a purulent discharge from
      the nose. Congestion may be relieved by antihistamines
      suitable for use on other species. As soon as the 2nd
      fever peak subsides, the ferret becomes more lively and
      soon recovers. Recovery itself differentiates influenza
      from the early stages of distemper and confers immunity
      for at least 5 weeks against the homologous influenza
      strain. Vaccination with attenuated live virus affords
      protection of a similar duration, but killed vaccine is
      ineffective.®

      Aleutian disease is usually a subclinical disease;
      however, affected ferrets may occasionally become hy-
      pergammaglobulinemic, become thin, pass black tarry
      feces, and eventually die.!° Ferrets with Aleutian dis-
      ease virus may carry it as long as 200 days. Because

      1156

      » Google

      the gross lesions are not prominent in ferrets, the disease
      must be diagnosed by use of the counter immunoelectro-
      phoresis test.11

      Ferrets are also susceptible to pseudorabies virus
      infection, although natural outbreaks of the disease have
      not been reported. In addition, infectious bovine rhino-
      tracheitis virus has been isolated from clinically normal
      ferrets (assumed to have been introduced in the diet
      through contaminated beef by-products).

      Bacterial Diseases—Ferrets are moderately suscep-
      tible to botulism (C botulinum) types A and B, and
      they are highly suspectible to botulism type C. Signs
      of muscular incoordination and stiffness appear in
      affected ferrets within 12 to 96 hours after eating con-
      taminated meat or poultry. They may salivate and,
      when picked up, will hang limp. The toxin paralyzes
      skeletal muscles, including the muscles of respiration,
      and the ferret will die of anoxia. There is no success-
      ful treatment, although the disease can be prevented by
      excluding from the diet food ingredients of unknown
      origin or by prophylactic use of toxoid.

      Avian, bovine, and human species of mycobacteria
      cause tuberculosis in ferrets. These species affect pri-
      marily the alimentary tract and abdominal lymph nodes.
      Outward signs may not appear until the late stages of
      active infection, when the animal becomes emaciated
      and may have paralysis of the adductor muscles of the
      pelvic limbs. Paralysis later affects all 4 limbs.'* Histo-
      logically, massive infiltrations of histiocytes in abdom-
      inal organs and intracellular acid-fast bacilli are diagnos-
      tic. Tuberculosis may be suspected if the animal gives
      a thermal response to the subcutaneous tuberculin test
      or if it has palpably enlarged mesenteric lymph nodes.

      Staphylococcal and streptococcal infections may
      cause abscesses arising from injuries to the mouth by
      bones in the diet or from bite wounds inflicted during
      the breeding season by aggressive males. Vulval infec-
      tions are common in the female during estrus. Strepto-
      coccus zooepidemicus causes pneumonia and metritis in
      this species, and Staphylococcus aureus causes mastitis.
      Treatment should follow guidelines established for cats
      and dogs. However, although ferrets can be treated
      with maximum recommended doses of penicillin, they
      should be given no more than 50 mg of streptomycin at
      12-hour intervals because it is extremely toxic at higher
      doses.

      Ferrets are undoubtedly also susceptible to many of
      the bacterial diseases that may affect mink, eg, anthrax,
      tularemia, and brucellosis, but natural outbreaks have
      not been reported. Normal animals are resistant to oral
      exposure of salmonella, but the organism can cause preg-
      nant females to abort. Ferrets may harbor and trans-
      mit Listeria monocytogenes without showing clinical
      signs of disease." ~

      Parasites and Fungi—Ferrets are susceptibie to j@-
      festation with Sarcoptes scabiet. The feet become swol-
      len, scabby, and if left untreated, clawless. Treatnient
      consists of cutting back the claws and removing the
      scabs after softening them in soapy water. Judicious
      application of sulfur ointment or lime and suliur dips
      and washes (30% to 32% calcium polysulfide) are effec-

      JAVMA, Vol 173, No. 9
      Ca

      tive, but all bedding must be burned and the cage
      disinfected to prevent reinfestation. Organophosphate
      compounds should be used only with caution, as levels
      safe for use in ferrets have not yet been established.

      Ringworm (Microsporum canis) has been reported
      in young ferrets and may be transmitted by cats.'!° Le-
      sions appear as large circumscribed areas of alopecia and
      inflammation on all parts of the body. The skin becomes
      thickened, red, and covered with scaling crusts. Al-
      though clinical signs often regress without treatment,
      affected ferrets may be treated orally with griseofulvin
      (25 mg/kg).

      Ferrets also may be infested with fleas (Ctenoceph-
      alides spp) and ear mites (Otodectes cynotis) and may
      be treated with rotenone or pyrethrin powders. They
      are also susceptible to coccidiosis, cryptococcosis,!* toxo-
      plasmosis,!’ actinomycosis,!* and various species of gas-
      trointestinal helminths. Guidelines for cats should be
      followed when treatment for any of these conditions is
      required.

      Miscellaneous Problems

      An enteritis of unknown cause appears frequently in
      warm weather. Acute cases include bloody diarrhea,
      anorexia, and death within 3 to 4 days. In chronic cases
      of enteritis, diarrhea may be intermittent and appetite
      normal, but the animal fails to digest its food, becomes
      emaciated, and usually dies within 1 month. For
      treatment of the disease, neomycin (10 to 20 mg/kg)
      or other products that are not absorbed by the gut
      should be used in the same dosage that would be pre-
      scribed for the cat.

      When fed ground chicken or turkey heads or other
      animal by-products, a ferret may get a small tracheal
      ring or blood vessel encircled and lodged around the base
      of the tongue. The ferret appears hungry but refuses
      to eat and may salivate and scratch at its mouth. The
      foreign body should be removed at once, to prevent
      necrosis of the tongue and death of the animal.

      Ferrets are also subject to hair loss, particularly if
      fed a diet containing an excessive amount (over 10%)
      of raw chicken eggs. Avidin, an enzyme in the egg
      white, causes a biotin deficiency that results in bilateral
      alopecia. On the other hand, we have observed that
      ferrets may lose their hair for unknown reasons, even
      when ringworm or external parasites are not involved.
      Such animals will usually respond to the addition of
      5% to 10% uncooked liver to the diet.

      Eclamptogenic toxemia, which may occur a few days
      before whelping, can result in the death of unborn kits
      as well as the dam. The only characteristic postmortem
      change is a yellow liver. The cause of the disease is
      not well understood, although it may be nutritional and
      can usually be prevented by the addition of uncooked
      fresh liver to the ration during pregnancy.

      Postparturient females may be afflicted with “nursing
      sickness”: signs include anorexia, weight loss, weakness,
      and muscular incoordination. Death follows a period of
      coma. The condition, which is usually first observed in
      late June, at about the time the young kits are weaned,
      is believed to result from a dietary salt insufficiency and

      November I, 1978

      Google

      may be corrected by the addition of 0.05% table salt
      to the feed from late May to mid-July.

      Under normal conditions, cannibalism is a minor
      problem in ferrets and can usually be prevented by
      providing a clean, dry, warm environment, a well-bal-
      anced diet, and easy access to water. Although canni-
      balism is most often pronounced in certain families or
      breeds and may therefore be an undesirable genetic
      trait, most dams and kits will normally consume a dead
      litter member.

      In some colonies, preweaning deaths in litters may
      approach 70%.1* Contributing causes include maternal
      lactating failure, maternal neglect or dietary insuffi-
      ciency, unclean or overheated nesting boxes that permit
      the kits to sweat and stick together, and viral or bac-
      terial infections. Although many of the kits die within
      the first 48 hours, later mortalities due to maternal
      lactating failure may be prevented if kits are given easy
      access to soft, solid food as early as 2 weeks of age.

      Litters of ferret kits are also subject to a variety of
      spontaneous congenital malformations that may be
      hereditary. These include anencephaly, cryptorchidism,
      absence of the tail or frontlimbs or hindlimbs, corneal
      hair growth, and eye opacity. Willis and Barrow? re-
      ported spontaneous neuroschisis or gastroschisis in about
      3% of their ferret kits. Posterior paralysis, generally ac-
      companied by lack of excretory control, may appear sud-
      denly in an otherwise healthy animal. It may be caused
      by hemivertebra or an inherited spinal weakness lead-
      ing to a broken back and is irreversible when spinal cord
      damage is severe.

      In addition, ferrets have been reported to suffer from
      lymphadenoma, diabetes mellitus,!® squamous cell car-
      cinoma, and an enzootic malignant granulomatosis of
      unknown cause. The latter has been reported to affect
      nearly 12% of certain ferret populations, attacking pri-
      marily animals less than 1 year old.?° It is characterized
      by loss of appetite and weight, listlessness, and upon
      necropsy, infiltrative granulomatous process in the mes-
      enteric lymph nodes and abdominal organs.

      References

      1. Threatened Wildlife of the United States. Publication 114,
      US Government Printing Office, Washington, DC, US Depart-
      ment of the Interior, Bureau of Sport Fisheries and Wildlife,
      1973, p 289.

      2. Willis LS, Barrow MV: The ferret (Mustela putorius
      furo L) as a laboratory animal. Lab Anim Sci 21:712-716, 1971.

      3. Doty BA, Jones CN, Doty LA: Learning-set formation
      by mink, ferrets, skunks, and cats. Science 155:1579-1580, 1967.

      4. Hammond J: The ferret: Some observations on photo-
      period and gonadal activity, and their role in seasonal pelt and
      bodyweight changes. Cambridge, W Heffer and Sons, Ltd, 1974,
      pp 3-11.

      5. Melby EC, Altman NH (ed): Handbook of Laboratory
      Animal Science. Cleveland, CRC Press Inc, 1974, vol 1, pp 108-
      109.

      6. Hammond J, Chesterman FC: The ferret. In The Uni-
      versities Federation for Animal Welfare Handbook on the Care
      and Management of Laboratory Animals. London, UFAW
      Churchill Livingstone, 1972, pp 345-362.

      7. Ott RL, Svehag SE, Burger D: Resistance to experimen-
      tal distemper in ferrets following the use of killed tissue vac-
      cine. Western Vet 6:107-111, 1959.

      8. Williams CSF: A Practical Guide to Laboratory Animals.
      St Louis, CV Mosby Co, 1976, pp 65-71.

      9. Potter CW, Oxford JS, Shore SL, et al: Immunity to in-
      fluenza in ferrets, I. Responses to live and killed virus. Br J
      Exp Pathol 53:153-167, 1972.

      10. Ohshima K, Shen DT, Henson JB, et al: Comparison

      1157
      of the lesions of Aleutian disease in mink and hypergamma-
      globulinemia in ferrets. Am J Vet Res 89:653-657, 1978.

      11. Cho HJ, Ingram DG: Antigen and antibody in Aleutian
      disease in mink, II. The reaction of antibody with the Aleutian
      disease agent using immunodiffusion and immunoelectrophoresis.
      Can J Comp Med 37:217-223, 1973.

      12. Porter DD, Larsen AE, Cox N:
      bovine rhinotracheitis virus from Mustelidae.
      1:112-113, 1975.

      13. Symmers WStC, Thompson APD: Observations on tu-
      berculosis in the ferret (Mustela furo L). J Comp Pathol 63:
      20-30, 1953.

      14. Morris JA, Norman MC: The isolation of Listeria mono-
      cytogenes from ferrets. J Bacteriol 59:313-314, 1950.

      15. Hagen KW, Gorham JR: Dermatomycoses in fur an-

      Isolation of infectious
      J Clin Microbiol

      1158

      imals: Chinchilla, ferret, mink and rabbit. Vet Med Small
      Anim Clin 67:438-48, 1972.

      16. Skulski G, Symmers WStC: Actinomycosis and torulosis
      in the ferret. J Comp Pathol 64:306-311, 1954.

      17. Lainson R: Symposium on _ toxoplasmosis, III. The
      demonstration of toxoplasm in animals with particular reference
      to members of the Mustelidae. Trans R Soc Trop Med Hyg
      51:111-117, 1957.

      18. Seamer J, Chesterman FC: A survey of disease in lab-
      oratory animals. Lab Anim 1:117-139, 1967.

      19. Carpenter JW, Novilla MN: Diabetes mellitus in a
      black-footed ferret. JAVMA 171:890-893, 1977.

      20. Momberg-Jorgensen HC: Enzootic malignant granulo-
      als in ferrets. Acta Pathol Microbiol Scand 29:297-306,
      1 "

      JAVMA, Vol 173, No. 9

    Attachments

    • PubMed entry
    • Ryland and Gorham - 1978 - The ferret and its diseases.pdf
  • The genetic legacy of the 19th‐century decline of the British polecat: evidence for extensive introgression from feral ferrets

    Item Type Journal Article
    Author M. Costa
    Author C. Fernandes
    Author J. D. S. Birks
    Author A. C. Kitchener
    Author M. Santos‐Reis
    Author M. W. Bruford
    Abstract In the 19th century, the British polecat suffered a demographic contraction, as a consequence of direct persecution, reaching its lowest population in the years that preceded the First World War. The polecat is now recovering and expanding throughout Britain, but introgressive hybridization with feral ferrets has been reported, which could be masking the true range of the polecat and introducing domestic genes into the species. We used a fragment of the mitochondrial DNA control region and 11 microsatellite loci to characterize the frequency and extent of hybridization and introgression between the two species and assess whether the 19th-century decline corresponded to a genetic bottleneck in the polecat. The proportion of admixture detected in the wild was high (31%) and hybrids were more frequently found outside Wales, suggesting that hybridization is more likely to occur along the eastern edge of the polecat’s range expansion. The patterns observed in the mitochondrial and nuclear DNA data show that introgression was mediated by crosses between male polecats and female ferrets, whose offspring backcrossed with polecats. No first-generation (F1) hybrids were identified, and the broad range of observed admixture proportions agrees with a scenario of past extensive hybridization between the two species. Using several different methods to investigate demographic history, we did not find consistent evidence for a genetic bottleneck in the British polecat, a result that could be interpreted as a consequence of hybridization with ferrets. Our results highlight the importance of the Welsh polecat population for the conservation and restoration of the genetic identity of the British polecat.
    Date 10/2013
    Language en
    Short Title The genetic legacy of the 19th‐century decline of the <span style="font-variant
    Library Catalog DOI.org (Crossref)
    URL https://onlinelibrary.wiley.com/doi/10.1111/mec.12456
    Accessed 5/21/2025, 4:10:43 PM
    Rights http://onlinelibrary.wiley.com/termsAndConditions#vor
    Volume 22
    Pages 5130-5147
    Publication Molecular Ecology
    DOI 10.1111/mec.12456
    Issue 20
    Journal Abbr Molecular Ecology
    ISSN 0962-1083, 1365-294X
    Date Added 5/21/2025, 4:10:43 PM
    Modified 10/21/2025, 5:10:00 PM

    Attachments

    • Costa et al. - 2013 - The genetic legacy of the 19th‐century decline of the Bspan.pdf
  • The History of Farm Foxes Undermines the Animal Domestication Syndrome

    Item Type Journal Article
    Author Kathryn A. Lord
    Author Greger Larson
    Author Raymond P. Coppinger
    Author Elinor K. Karlsson
    Date 2020-02-01
    Language English
    Library Catalog www.cell.com
    URL https://www.cell.com/trends/ecology-evolution/abstract/S0169-5347(19)30302-7
    Accessed 7/14/2025, 4:21:58 PM
    Extra Publisher: Elsevier PMID: 31810775
    Volume 35
    Pages 125-136
    Publication Trends in Ecology & Evolution
    DOI 10.1016/j.tree.2019.10.011
    Issue 2
    Journal Abbr Trends in Ecology & Evolution
    ISSN 0169-5347
    Date Added 7/14/2025, 4:21:58 PM
    Modified 7/14/2025, 4:21:58 PM

    Tags:

    • Anthropocene
    • domestic animals
    • domestication syndrome
    • behavioral selection
    • neural crest
    • pleiotropic effects
    • silver fox

    Attachments

    • Full Text PDF
    • PubMed entry

    Related

    • Straw Foxes: Domestication Syndrome Evaluation Comes Up Short
  • Urban animals—domestic, stray, and wild: notes from a bear repopulation project in the Alps

    Item Type Journal Article
    Author Andrea Mubi Brighenti
    Author Andrea Pavoni
    Abstract This piece explores “domesticity” as a social territory defined by its relationship with the conceptual and ecological space of “the wild,” and asks whether these spaces stand in opposition to each other or more subtle relations of co-implication are at play. As we look into the domestic and the wild, a conceptual map of notions emerges, including the public, the common, the civilized, and the barbarian. The paper suggests the domestic and the wild constitute two semiotic-ecological domains constantly stretching into each other without any stable or even clear boundary line, and it elaborates on a series of corollaries for studying non-human animals in urban contexts. As an illustrative case study, we follow the story of Daniza, a wild brown bear introduced in the Brenta Natural Park on the Italian Alps in the 2000s. Declared a “dangerous animal,” Daniza was accidentally, and controversially, killed by the public authorities in 2014.
    Date 2018-12-07
    Language en
    Short Title Urban animals—domestic, stray, and wild
    Library Catalog DOI.org (Crossref)
    URL https://brill.com/view/journals/soan/26/6/article-p576_2.xml
    Accessed 3/11/2020, 12:33:00 PM
    Volume 26
    Pages 576-597
    Publication Society & Animals
    DOI 10.1163/15685306-12341580
    Issue 6
    Journal Abbr Soc. Animals
    ISSN 1063-1119, 1568-5306
    Date Added 3/11/2020, 12:33:00 PM
    Modified 6/14/2025, 9:25:23 AM

    Attachments

    • Brighenti and Pavoni - 2018 - Urban animals—domestic, stray, and wild notes fro.pdf
  • Using canine width to determine age in the black-footed ferret Mustela nigripes

    Item Type Journal Article
    Author Rachel M. Santymire
    Author Samantha M. Wisely
    Author Travis M. Livieri
    Author J. Howard
    Date 2012
    Library Catalog Google Scholar
    Volume 46
    Pages 17–21
    Publication Small Carnivore Conservation
    Date Added 10/27/2017, 9:22:15 AM
    Modified 5/26/2025, 10:31:55 PM

    Attachments

    • Santymire et al. - 2012 - Using canine width to determine age in the black-f.pdf
  • Zoonoses of Ferrets, Hedgehogs, and Sugar Gliders

    Item Type Journal Article
    Author Charly Pignon
    Author Jörg Mayer
    Date 9/2011
    Language en
    Library Catalog DOI.org (Crossref)
    URL https://linkinghub.elsevier.com/retrieve/pii/S1094919411000442
    Accessed 5/21/2025, 4:31:43 PM
    Rights https://www.elsevier.com/tdm/userlicense/1.0/
    Volume 14
    Pages 533-549
    Publication Veterinary Clinics of North America: Exotic Animal Practice
    DOI 10.1016/j.cvex.2011.05.004
    Issue 3
    Journal Abbr Veterinary Clinics of North America: Exotic Animal Practice
    ISSN 10949194
    Date Added 7/14/2023, 12:11:14 PM
    Modified 5/26/2025, 10:31:41 PM

    Attachments

    • Pignon and Mayer - 2011 - Zoonoses of Ferrets, Hedgehogs, and Sugar Gliders.pdf